Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)
Author
Gussmann S. M.
Author
Scholtz, C. H.
text
Journal of Natural History
2000
2000-12-31
34
1045
1123
journal article
10.1080/00222930050020122
1464-5262
4747979
Meridiobolbus faustus
(PeÂringuey)
(®gures 9a±c, 15a, b, 16a±c, 18a,
19, 24e
)
Bolboceras faustum
PeÂringuey, 1908: 639
;
Boucomont 1912: 9
;
Paulian 1941: 26
.
Meridiobolbus faustus
(PeÂringuey)
:
Krikken 1984: 38
.
Description
Body length
.
11.4±17.7 mm
.
Colour
. Mid- to dark brown.
Head
.
Outer margin of mandible
rounded but anteriorly more strongly curved inward (®gure 15a).
Labrum
(®gure 15a) declivous anteriad and posteriad from feebly developed, slightly arcuate transverse carina; anterior margin distinctly sinuate.
Clypeus
(®gures 15a, 18a) with strongly elevated transverse anterior carina, in dorsal view medially bowed; with well-de®ned, short diagonal, posteriorly steeply inclined carina, the latter connecting the antero-lateral marginal angle and the respective endpoint of the transverse anterior carina; with distinct lateral carina from endpoint of anterior carina extending to above antennal insertion, from there extending posteriorly in a slightly curved line past eye-canthus on frons reaching posterior end of frons.
Frons
(®gures 15a, 18a) with bituberculate, strongly elevated transverse carina, positioned between eye-canthi.
Pronotum
(®gure 18a). With four equally developed, tuberculate swellings in anterior third; areas between and anteriad to tubercles slightly concave; with feebly indented midline, not reaching anterior declivous part or posterior pronotal margin; with ®ne punctures in posterior half, interspersed by some large punctures, especially along midline; ®ne punctation increasingly replaced by large, partly conūent punctures anteriorly and towards sides; tubercles with ®ne punctures only.
Scutellum
(®gure 15b). Basal width about 1.44Ölength; with large, shallow punctures, particularly in the middle.
Elytron
(®gure 15b). Striae strongly impressed, their punctures small and separated by three to four puncture diameters; intervals with double punctation (approximately four to six ®ne and 10 minute punctures between two striae).
Legs
.
Protibia
®ve-dentate.
Metafemur
with punctation and setation of ventral side as in
Bolboceratex
(®gure 24e).
Metatibia
in lateral view elongate, only slightly broadening apically; with subapical carina strongly bilobed; with toothlike projections arranged in two rows above subapical carina.
Male genitalia
. Aedeagus as in ®gure 16a±c.
Distribution
(®gure 19). Western
Botswana
, southern
Zimbabwe
and northeastern
South Africa
.
Type material examined
.
LECTOTYPE
(here designated):`
Bolboceras faustum Typ.
Py’
[handwriting PeÂringuey]/`
Plumtre
e [
20.30S
27.50E
],
Rhodesia
, 28/1/03, to light’ /`SAM’ /`
Type
SAM/
Ent
5078’ [black print and handwriting on red label] (
SAMC
).
Additional material examined
(
117 specimens
).
Botswana
:
2x,
Botsw.
,
Mahalapye
[
23.04S
26.50E
],
26.iii.1977
,
A. M. Camerith
(
TMSA
)
;
1x,
Botswana
,
110 km
S
Francistown
[
22.03S
27.16E
],
12.i.1978
,
Holm
,
Jakobs
,
Kirsten
,
Scholtz
(
UPSA
, 97003)
.
South Africa
:
1x,
Transv.
,
Pietersburg
[
23.53S
29.24E
], ... [illegible] (
SAMC
)
;
1x,
Pietersburg
,
21.iv.1961
,
A. Spies
(
TMSA
)
; 1x, Pietersburg, Janse
(SAMC);
1x,
Pietersburg
, ... [illegible] (
SANC
)
;
1x,
Orange Free State
,
Distr.Parys
,
Farm Abel
,
26.52S
27.37E
,
27.xii.1991
,
C. H. Scholtz
(
UPSA
)
;
1x,
Naboomspruit
[
24.32S
28.36E
], W.P.,
4.ii.1939
, A.R.I. (
UPSA
)
;
1x,
Pretoria
[
25.43S
28.11E
],
i.1961
,
F.J.Kleynhans
(
UPSA
)
;
1x,
Pretoria
,
5.ii.1910
,
A. J. T. Janse
(
TMSA
)
;
1, l
Pretoria
,
xii.1923
,
G. v. Son
(
TMSA
)
;
1x,
Pretoria
,
15.v.1967
, B.S. (
UPSA
)
;
1, l
Waterkloof
,
Pret. Distr.
,
xi.1903
,
F. Noome
(
TMSA
)
; 1, l M.A.,
ii.1971
, Pret.
(UPSA); 1, l N.R.,
15.iv.1972
,
Pret.
(
UPSA
); 1x,
North
Rand
,
Transvaal
,
ii.1944
,
G.Kobrow
(
TMSA
); 1x,
Johannesburg
[
26.12S
28.05E
],
6000 ft.
,
i.1896
, J. P.
Cregoe
(
MNHN
); 1x, same data but
:
ii.1896
(
SAMC
); 1x,
Johannesburg
,
iii.1937
, G.
Kobrow
(
TMSA
); 1x, same data but
:
i.1907
(
TMSA
); 1x, same data but
:
xii.1906
(
TMSA
); 1x, same data but
:
15.xii.1901
, at light (
TMSA
); 1x, same data but
:
xii.1902
, at light (
TMSA
);
1m
,
Jhb
,
Frankenwald
,
20.ii.1970
,
R. H. Taylor
(
UPSA
); 1x, C.C.,
Portuani
[not traced], 1907,
H. Bazeley
(
SAMC
); 1x,
Tvl.
,
Rivonia
,
9.ii.1967
,
M. J. Russell
,
Brit. Mus.
1972-258 (
BMNH
); 1, l
Vryburg
[
26.57S
24.44E
],
Louis
[illegible], 04 (
SAMC
); 2x,
Transvaal
,
Warmbad
[
24.53S
28.18E
] (
ZMHB
); 1, l
Warmbad
,
Tvl
.,
29.xii.1988
,
A.Joubert
(
AJCS
); 1, l S.A.,
Rietvlei Dam
,
25.44S
28.16E
,
1.iii.1989
, leg.
K. T. Language
(
UPSA
); 1, l
Transvaal
,
Waterval Farm
,
25.35S
28.16E
,
2.ii.1991
, light, leg.
R. Dixon
(
TMSA
); 1x, same data but
:
24.xii.1992
(
TMSA
); 1, l
Nelspruit
[
25.32S
30.54E
]
,
24.xi.1988
,
D.J.Kotze
(
UPSA
); 1x,
Modderpoort
[
29.07S
27.28E
]
,
18.xii.1924
,
A.J.T.Janse
(
TMSA
); 1x,
Nylstroom
,
24.39S
28.25E
,
2.iii.1990
,
F.P.Schoeman
(
UPSA
); 2x,
Transv.
,
Rustenburg
[
25.40S
27.15E
]
,
8-12.xii.1952
,
D. W. Rorke
(
TMSA
); 1x,
Rustenburg
[
25.40S
27.15E
]
,
27.x.1966
, A. L.
Capener
(
SANC
); 3x,
North-West
,
Rustenburg Nat. Res.
,
25.42S
27.13E
,
28.ii.1998
, leg.
C. H. Scholtz
, at light (
UPSA
); 1x,
Noord-Wes
,
Rustenburg
,
25.40S
27.11E
,
28.ii.1998
, leg.
S. Pieterse
(
UPSA
); 1x,
Rustenburg Natuurreservaat
,
25.42S
27.14E
,
28.ii.1998
, leg.
E. A. M. Booyens
(
UPSA
); 1x,
Rustenburg
,
25.43S
27.12E
,
28.ii.1998
, R.
Olivier
(
UPSA
); 1x,
Noordwes
,
Rustenberg
N.R.
,
25.43S
27.12E
,
28.ii.1998
, J. A.
Maatens
(
UPSA
);
1m
,
North-West
,
Rustenberg N
/R,
25.13S
27.11E
,
22.ii.1997
,
B. Botha
(
UPSA
); 1, l
Transv.
,
Cullinan Distr.
,
H.D. Khotso Farm
[not traced], i
.&
ii.1980
, leg.
E. Voight
(
TMSA
); 1x,
Kimberley
[
28.45S
24.46E
] (
ZMHB
); 1x,
Kimberley
, 1920,
J.H.Power
(
SAMC
); 1x,
Middelburg
[Tvl?
25.47S
29.28E
], 1907,
Vits
(
ISNB
); 1x,
Middelburg
,
10.iii.1910
,
G.Hewilt
(
TMSA
); 1x,
Transv.
,
Tzaneen
63km
S of [
ca
SE 24 30 Da]
,
29.xii.1977
,
J.Boomker
(
UPSA
);
1m
,
Pretoria
,
Verwoerdburg
[
25.50S
28.13E
]
,
19.ii.1991
, wild olive tree, leg.
L.D.Jemings
(
TMSA
); 1x,
Barberton
[
25.47S
31.03E
]
,
xii.1940
,
W. G. Kobrow
(
TMSA
); 1x,
Potchefstroom
[
26.43S
27.06E
]
T. Ayres
(
TMSA
); 3x,
Bultfontein
[
28.12S
26.09E
]
,
8.i.1907
(
TMSA
); 1x, same data (
HAHC
); 5x,
Gri
n
Mine
[near Leydsdorp,
23.59S
30.31E
]
,
i.1915
,
H.G.Breijer
(
TMSA
); 2x,
Zoutpan
[
25.24S
28.06E
],
Pta
,
9.xii.1926
,
G. van Son
(
TMSA
); 1x,
Pretoria Distr.
,
Soutpan
,
25.24S
28.06E
,
9.xii.1987
,
M.W.Mansell
(
SANC
); 1x,
Tweefontein
[
NE Pretoria
?]
,
13.i.1907
(
TMSA
); 1x,
Vaal Rivier
,
Bargage
[not traced]
,
10.xii.1931
,
G.v.Son
(
TMSA
); 2x,
Moor
[d] drift, [O.
F.S.
,
24.17S
28.58E
]
x.1909
,
C. J. Swierstra
(
TMSA
); 2x,
Grootdraai
[not traced],
Olifants Riv., N.E
.
Tvl
.,
x.1927
,
H.Lang
(
TMSA
); 1x,
Leydsdorp
[
23.59S
30.32E
]
,
16.x.1907
,
J. Maughton
(
TMSA
); 1x,
Tvl.
,
Bundu Inn
, nr.
Verena
[Verena:
25.29S
29.01E
]
,
5.xi.1973
, at camp site,
E. Breytenbach
(
TMSA
); 1x,
Ndumu
[
26.53S
32.16E
],
Zulul
.,
xii.1960
, van
Son
(
TMSA
); 1x, BuOEelspoort [
25.48S
27.29E
]
,
17.xii.1924
,
A. J. T. Janse
(
TMSA
); 1, l
Pretoria
, UV-Lichtfalle,
Farm Roodeplaat
[
ca
25.38S
28.23E
]
,
vii.1961
,
P. Neubecker
(
HNHM
); 1x,
Thabazimbi
,
Tweeloopf
50 km
NW, SE 24 27
Ac
,
26.iii.1989
,
E. Holm
,
E. Marais
(
UPSA
); 1x,
Transvaal
,
Argent
[
26.03S
28.49E
], 7±
10.xii.1968
,
A. L. Capener
(
HAHC
)
;
1m
,
Roodepoort
,
26.11S
27.51E
,
1.iv.1997
,
H. Laity
(
UPSA
)
;
1m
,
North-West
,
Dikhololo
,
25.22S
27.34E
,
14.ii.1997
,
H. Gray
(
UPSA
)
;
1, l
Transvaal
,
Frm.
:
Rhenosterpoort
,
25.43S
28.56E
,
15.ii.1980
, leg.
Schulze
(
TMSA
)
;
1, l
Transvaal
,
Waterval Farm
,
25.35S
28.16E
,
2.ii.1993
, at light, leg.
R. Dixon
(
TMSA
)
;
1x, same data but:
30.xii.1976
, E-Y: 1319, on gas-lamp light, leg.
Proz.
-
Schulze
(
TMSA
)
;
1x,
S. Afr.
,
Kruger Nat. Park
,
Skukuza Rest Camp
,
24.59S
31.36E
, E-Y: 3120,
UV light trap
, leg. Endroedy-
Younga
(
TMSA
)
;
1x,
Skukuza
,
E.Tvl.
,
5.x.1971
, T.B. &
A.P.
(
SANC
)
; 1, l4x (SANC), 1x (HAHC),
Skukuza
[
25.00S
31.35E
], KNP,
1.xii.1971
,
H. D. Catling
(
SANC
)
;
1x,
Hennops River
,
v.1967
, V. H.
Wahl
(
HAHC
)
;
1, l5x,
Groenvlei
[
24.15S
27.02E
],
3.i.1907
(
TMSA
)
;
3x,
Tvl
,
Hoedspruit
,
Vienna Game Farm
,
24.17S
30.58E
,
500 m
, 17±
18.i.1991
,
B. Grobbelaar
(
SANC
)
;
1x, NWP,
Veekraal
(
28.20S
27.44E
),
23.iii.1996
,
J. Marshall
(
UPSA
)
;
2x,
Fount. Gr.
[not traced],
ii.1901
,
Jutrzencka
(
SANC
)
;
1x,
Rietf.
[not traced],
31.i.1907
(
TMSA
)
; 4x (TMSA), 1x (SAMC),
no locality data.
Zimbabwe
:
3x (
ISNB
)
, 1x (SANC), 1x (SAMC),
Plumtree
[
20.30S
27.50E
],
Rhod.
,
8.i.1907
(
ISNB
)
;
1x, same data but:
2.i.1907
(
SAMC
)
;
1x,
S. Rhod.
,
Insiza
[Nsiza
19.44S
29.12E
],
i.1919
,
G. French
(
SAMC
)
;
1x, same data but:
i.1949
, G.
French
(
SAMC
)
;
1x,
Hillside
[
20.10S
28.35E
],
S.Rh.
,
28.ii.1923
,
Swinburne
&
Stevenson
(
TMSA
)
;
1x,
S. Rhodesia
,
Empandeni
[Empandine,
20.43S
27.55E
], 1911,
Rev. J.
O’
Neil
(
SAMC
)
;
1x,
Bulawayo
[
20.10S
28.43E
],
27.xii.1924
, R. H. R.
Stevenson
(
TMSA
)
;
1x,
S. Rhodesia
,
Khami
[
20.18S
28.27E
]/
Bulawayo
(aÁ la lumieÁre),
10.i.1954
,
Mr Cooke
(
MRAC
)
.
Comments
. Intraspeci®c variability between major and minor males and between males and females of
Meridiobolbus faustus
(PeÂringuey)
is very small. The bituberculate transverse carina on the frons immediately distinguishes
Meridiobolbus faustus
from its congeneric species.