A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Author Wild, A. L. text Zootaxa 2007 1622 1 55 http://www.antbase.org/ants/publications/21367/21367.pdf journal article 21367 asper (Mayr 1887). Alto Parana , Canindeyu , Paraguari (ALWC, IFML, INBP, MZSP).