Checklist of the marine and estuarine fishes of Madang District, Papua New Guinea, western Pacific Ocean, with 820 new records Author Fricke, Ronald Author Allen, Gerald R. Author Andréfouët, Serge Author Chen, Wei-Jen Author Hamel, Mélanie A. Author Laboute, Pierre Author Mana, Ralph Author Hui, Tan Heok Author Uyeno, Daisuke text Zootaxa 2014 3832 1 1 247 journal article 45252 10.11646/zootaxa.3832.1.1 8646401a-8028-4795-a6d6-1f9d75607e41 1175-5326 250559 BAB612A4-03DB-4958-BEB1-92DE278FBB90 Kyphosidae Kyphosus cinerascens (Forsskål in Niebuhr, 1775) —Topsail chub STATUS AT MADANG. New record from Madang, based on specimens photographed by A. Berberian. DISTRIBUTION AND HABITAT. Red Sea, East Africa, Madagascar and Mascarenes east to Hawaiian, Line and Easter islands, north to southern Japan , south to Western Australia , Lord Howe, Norfolk and Austral islands. Found in groups, over hard bottoms covered with algae, on exposed reef flats and seaward coral reefs, 0– 24 m . Marine. Kyphosus vaigiensis (Quoy & Gaimard, 1825) —Lowfin chub STATUS AT MADANG. New record from Madang, based on specimens photographed at St. CTR 13. DISTRIBUTION AND HABITAT. Red Sea, East and South Africa , Madagascar and Réunion east to Hawaiian, Line and Marquesas islands, north to central Japan , south to Western Australia , New South Wales ( Australia ), northern New Zealand , Tonga and Rapa. Found in groups, over hard bottoms covered with algae, on exposed reef flats and seaward coral reefs, 0– 24 m . Marine. Chaetodontidae Chaetodon auriga Forsskål in Niebuhr, 1775 —Threadfin butterflyfish STATUS AT MADANG. First record from Madang as Linophora setifer by Munro (1958: 216) . Specimens photographed at St. CTR 24, PR 228. DISTRIBUTION AND HABITAT. Red Sea, East and South Africa east to Hawaiian and northern Line islands and Pitcairn Group, north to southern Japan , south to northern New Zealand , and Rapa. Found in live and dead coral reef and lagoon habitats, also on rocky reefs, 0– 35 m . Marine. Chaetodon baronessa Cuvier, 1829 —Eastern triangular butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 3, CTR 0 4, CTR 0 5, CTR 0 7, CTR 0 8, CTR 0 9, CTR 10, CTR 12, CTR 13, CTR 14, CTR 15, CTR 16, CTR 19, CTR 20, CTR 21, CTR 22, CTR 23, CTR 24, CTR 25, CTR 26, CTR 27, CTR 30, CTR 31, CTR 32, CTR 35, PR 180; a specimen collected by W.-J. Chen in Madang Bay. AMS, CAS and NTUM material (AMS I.19712- 022; CAS 65712, 65173, 65174, 65175, 65176, 65177; NTUM 10241). DISTRIBUTION AND HABITAT. Christmas and Cocos-Keeling islands and Indonesia east to Fiji , north to southern Japan and Ogasawara Islands, south to New South Wales ( Australia ) and New Caledonia . Living in pairs around Acropora corals, 1– 20 m . Marine. Chaetodon bennetti Cuvier in Cuvier & Valenciennes, 1831 —Bennett's butterflyfish STATUS AT MADANG. New record from Madang, based on specimens photographed at St. CTR 16, and by P. Laboute on 28 Nov. 2012 . DISTRIBUTION AND HABITAT. East Africa east to Caroline Islands and Tuamotu Archipelago, north to southern Japan , south to northern Australia , Lord Howe Island and Rapa. Solitary or in pairs, in coral-rich areas of lagoon or seaward coral reefs, 1– 30 m . Marine. Chaetodon burgessi Allen & Starck, 1973 —Burgess' butterflyfish STATUS AT MADANG. New record from Madang, based on a specimen photographed by A. Berberian. DISTRIBUTION AND HABITAT. Eastern Indonesia east to Caroline Islands and Tonga , north to Philippines , south to Solomon Islands . Solitary or in pairs, on steep drop-offs of seaward reefs, 20– 80 m . Marine. Chaetodon citrinellus Cuvier [ex Broussonet] in Cuvier & Valenciennes, 1831 —Speckled butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 3, CTR 0 4, CTR 0 5, CTR 0 6, CTR 0 7, CTR 0 8, CTR 0 9, CTR 11, CTR 14, CTR 15, CTR 22, CTR 23, CTR 25, CTR 28; specimens collected by W.-J. Chen at Madang Resort. CAS and NTUM material (CAS 65178, 65179: NTUM 10278). DISTRIBUTION AND HABITAT. East Africa east to Hawaiian and Tuamotu islands, north to southern Japan and Ogasawara Islands, south to northern Australia , Lord Howe and Norfolk islands. Solitary, in pairs or in small groups, on reef flats and in seaward coral reefs, 1– 36 m . Marine. Chaetodon ephippium Cuvier in Cuvier & Valenciennes, 1831 —Saddled butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 4, CTR 16, CTR 30, CTR 31. AMS material (AMS I.16748-010, I.17086-018). DISTRIBUTION AND HABITAT. India and Sri Lanka east to Hawaiian and Tuamotu islands, north to southern Japan and Ogasawara Islans, south to northern Australia and Lord Howe Island . Solitary or in pairs, in coral-rich areas of lagoon and seaward coral reefs, 0– 30 m . Marine. Chaetodon kleinii Bloch, 1790 —Klein's butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 3, CTR 0 4, CTR 0 5, CTR 0 7, CTR 0 8, CTR 0 9, CTR 10, CTR 11, CTR 13, CTR 14, CTR 15, CTR 16, CTR 19, CTR 20, CTR 21, CTR 22, CTR 23, CTR 24, CTR 25, CTR 27, CTR 28, CTR 30, CTR 31, CTR 32, CTR 33, CTR 35, PR 180; a specimen collected by W.-J. Chen on 11 Dec. 2012 . AMS, CAS, NTUM and WAM material (AMS I.17085-003; CAS 65180, 65181, 65182, 65183, 65184, 65185; NTUM 10279; WAM P.30350-026). DISTRIBUTION AND HABITAT. Red Sea, East and South Africa , Madagascar and Mascarenes east to Hawaiian, Line and Marquesas islands, north to southern Japan , south to Kimberleys (Western Australia ), Lord Howe Island and Tonga . Found in lagoon and seaward coral and rocky reefs, 2– 61 m . Marine. Chaetodon lineolatus Cuvier [ex Quoy & Gaimard ] in Cuvier & Valenciennes, 1831 —Lined butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed by G. R. Allen.—DISTRIBUTION AND HABITAT, Red Sea, East Africa and Madagascar . Mascarenes east to Hawaiian, Line and Marquesas islands, north to southern Japan , south to Western Australia , Lord Howe, Norfolk and Gambier islands. Found in lagoon and seaward coral reefs, 0– 171 m . Marine. Chaetodon lunula (Lacepède [ex Commerson ], 1802) —Raccoon butterflyfish STATUS AT MADANG. New record from Madang, based on specimens photographed by P. Laboute on 13 Dec. 2012 . DISTRIBUTION AND HABITAT. East Africa, Madagascar and Mascarenes east to Hawaiian and Line islands and Pitcairn Group, north to southern Japan , south to Western Australia , New South Wales ( Australia ), Lord Howe and Norfolk islands, and Rapa. Found on reef flats in lagoon and seaward coral and rocky reefs, including tidal pools, 0– 30 m . Marine. Chaetodon lunulatus Quoy & Gaimard, 1825 —Redfin butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 2, CTR 0 3, CTR 0 4, CTR 0 5, CTR 0 6, CTR 0 7, CTR 0 8, CTR 0 9, CTR 10, CTR 11, CTR 12, CTR 13, CTR 14, CTR 15, CTR 16, CTR 16B, CTR 17, CTR 19, CTR 20, CTR 21, CTR 22, CTR 23, CTR 24, CTR 25, CTR 26, CTR 27, CTR 31, CTR 32, CTR 35, PR 113; specimens collected by W.-J. Chen in Madang Bay. AMS, CAS, NTUM, ROM and WAM material (AMS I.16671-009, I.17088-007, I.17090-009; CAS 65188, 65189, 65190, 65191, 65192, 65193, 65194, 65195, 65196, 65197, 65198; NTUM 10240; ROM 67004; WAM P.29624- 002). DISTRIBUTION AND HABITAT. Malaysia and Indonesia east to Hawaiian, Line and Tuamotu islands, north to southern Japan , south to northern Australia , Lord Howe and Norfolk islands. Found in coral-rich areas of coral reefs, 0– 20 m . Marine. Chaetodon melannotus Bloch & Schneider, 1801 —Black-backed butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 2, CTR 0 6, CTR 11, PR 228. AMS material (AMS I.16746-007, I.17083-005). DISTRIBUTION AND HABITAT. Red Sea and East Africa, Madagascar and Mascarenes east to Marshall Islands and Tonga , north to southern Japan , south to northern Australia , Lord Howe and Norfolk islands. Found in coralrich areas of reef flats, lagoon and seaward reefs, 0– 20 m . Marine. Chaetodon meyeri Bloch & Schneider, 1801 —Meyer's butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed by G. R. Allen. DISTRIBUTION AND HABITAT. East Africa and Réunion east to Caroline, Line and Phoenix islands , north to Ryukyu Islands, south to Queensland ( Australia ), New Caledonia and Tonga . Found in coral-rich areas of lagoon and seaward reefs, 2– 25 m . Marine. Chaetodon ocellicaudus Cuvier in Cuvier & Valenciennes, 1831 —Spot-tail butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 5, CTR 0 6, CTR 0 7, CTR 0 8, CTR 0 9, CTR 11, CTR 12, CTR 14, CTR 15, CTR 19, CTR 21, CTR 22, CTR 23, CTR 24, CTR 26, CTR 28. DISTRIBUTION AND HABITAT. Malaysia and Indonesia east to Palau and Solomon Islands , north to Philippines , south to Queensland ( Australia ). Found in coral-rich areas of reef flats, lagoon and seaward coral reefs, 3– 50 m . Marine. Chaetodon octofasciatus Bloch, 1787 —Eight-banded butterflyfish ( Fig. 10 ) STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 5, CTR 0 8, CTR 12, CTR 26, CTR 36. CAS material (CAS 65186). DISTRIBUTION AND HABITAT. Sri Lanka east to Philippines , north to southern Japan , south to New Guinea and Solomon Islands . Found among branching corals, in coral-rich areas of sheltered coastal and lagoon reefs, 3– 20 m . Marine. Chaetodon ornatissimus Cuvier [ex Solander] in Cuvier & Valenciennes, 1831 —Ornate butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 3, CTR 0 9, CTR 10, CTR 14, CTR 16, CTR 22, CTR 24, CTR 28, CTR 32, PR 113. DISTRIBUTION AND HABITAT. Maldives east to Hawaiian and Line islands and Pitcairn Group, north to southern Japan and Ogasawara Islands, south to Lord Howe Island . Found in coral-rich areas in lagoon and seaward coral reefs, 1– 36 m . Marine. Chaetodon oxycephalus Bleeker, 1853 —Spot-nape butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 3, CTR 0 9, CTR 31. DISTRIBUTION AND HABITAT. Maldives and Chagos Archipelago east to Pohnpei ( Caroline Islands ) and Solomon Islands , north to Philippines , south to Queensland ( Australia ). Found in coral-rich areas in coastal and seaward coral reefs, 10– 40 m . Marine. Chaetodon pelewensis Kner, 1868 —Dot-and-dash butterflyfish STATUS AT MADANG. New record from Madang, based on specimens photographed at St. CTR 28. DISTRIBUTION AND HABITAT. Eastern Australia east to Easter Island , north to New Guinea , south to Lord Howe and Norfolk islands. Usually found in pairs, in seaward coral reefs, 1– 30 m . Marine. FIGURE 10 . Chaetodon octofasciatus (Chaetodontidae) , Madang Lagoon, P. Laboute, 8 Dec. 2012. Chaetodon plebeius Cuvier [ex Broussonet] in Cuvier & Valenciennes, 1831 —Bluespot butterflyfish STATUS AT MADANG. New record from Madang, based on specimens collected at St. CTR 17, CTR 26. AMS and MNHN material (AMS I.16746-005; MNHN uncat.). DISTRIBUTION AND HABITAT. Western Australia ; Queensland ( Australia ) and Philippines east to Fiji , north to southern Japan and Ogasawara Islands, south to Lord Howe and Norfolk islands. Solitary or in pairs, in coastal, lagoon, and seaward coral reefs, 0– 10 m . Marine. Chaetodon punctatofasciatus Cuvier in Cuvier & Valenciennes, 1831 —Spot-banded butterflyfish STATUS AT MADANG. New record from Madang, based on specimens photographed at St. CTR 20. DISTRIBUTION AND HABITAT. Indonesia and Malaysia east to Line Islands, north to Ryukyu and Ogasawara islands, south to northern Australia . Usually reefs, in lagoon reefs and outer slopes of seaward reefs, 5– 45 m . Marine. Chaetodon rafflesii Anonymous [Bennett], 1830 —Latticed butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 2, CTR 0 3, CTR 0 4, CTR 0 6, CTR 0 7, CTR 11, CTR 13, CTR 14, CTR 15, CTR 20, CTR 22, CTR 23, CTR 26, CTR 27, CTR 31, CTR 32. AMS and CAS material (AMS I.17090-040; CAS 65187). DISTRIBUTION AND HABITAT. Sri Lanka east to Tuamotu Archipelago, north to Ryukyu Islands, south to northern Australia and New Caledonia . Solitary or in pairs, usually in coral-rich areas of coastal, lagoon and seaward coral reefs, 1– 15 m . Marine. Chaetodon semeion Bleeker, 1855 —Dotted butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed by G. R. Allen. DISTRIBUTION AND HABITAT. Sri Lanka east to Line and Tuamotu islands, north to Ryukyu Islands, south to Queensland ( Australia ) and New Caledonia . Usually in pairs, in coral-rich areas of lagoon reefs and outer slopes of seaward reefs, 2– 30 m . Marine. Chaetodon speculum Cuvier [ex Kuhl & Hasselt] in Cuvier & Valenciennes, 1831 —Oval-spot butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 30, CTR 33. DISTRIBUTION AND HABITAT. Indonesia east to Yap ( Caroline Islands ) and Tonga , north to southern Japan and Ogasawara Islands, south to northern Australia and New Caledonia . Usually solitary, occasionally in pairs, in coral-rich areas of lagoon and seaward coral reefs, 1– 30 m . Marine. Chaetodon trifascialis Quoy & Gaimard, 1825 —Chevroned butterflyfish ( Fig. 11 ) STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 9, CTR 10, CTR 14, CTR 15, CTR 20, CTR 22, CTR 25, CTR 28, CTR 30, CTR 32, PR 79. DISTRIBUTION AND HABITAT. Red Sea, East and South Africa east to northern Line and Tuamotu islands, north to southern Japan , south to Exmouth Gulf (Western Australia ), New South Wales ( Australia ), Lord Howe and Norfolk islands and Rapa. Found in coral-rich areas of shallow lagoon and semi-protected seaward coral reefs, 0– 30 m . Marine. Chaetodon ulietensis Cuvier in Cuvier & Valenciennes, 1831 —Pacific double-saddle butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 6, CTR 12, CTR 23, CTR 25, CTR 34. DISTRIBUTION AND HABITAT. Cocos-Keeling and Christmas islands (eastern Indian Ocean) east to Hawaiian and Line islands and Pitcairn Group, north to southern Japan and Ogasawara Islands, south to northern Australia and Lord Howe Island . Found solitary, in pairs or in groups, in coral-rich areas of lagoon and seaward coral reefs, 2– 30 m . Marine. Chaetodon unimaculatus Bloch, 1787 —Teardrop butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 10, CTR 13, CTR 16, CTR 20, CTR 25, CTR 28, CTR 32. DISTRIBUTION AND HABITAT. Cocos-Keeling and Christmas islands (eastern Indian Ocean) east to Hawaiian and Easter islands, north to southern Japan and Ogasawara Islands, south to northern Australia and Lord Howe Island . Solitary or in small groups, in lagoon and seaward coral reefs, 0– 60 m . Marine. Chaetodon vagabundus Linnaeus, 1758 —Vagabond butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 2, CTR 0 3, CTR 0 4, CTR 0 6, CTR 0 7, CTR 10, CTR 11, CTR 12, CTR 14, CTR 15, CTR 16B, CTR 17, CTR 18, CTR 21, CTR 22, CTR 23, CTR 24, CTR 26, CTR 30, CTR 31, CTR 35, PR 210. AMS material (AMS I.17081-006, I.17082-010). DISTRIBUTION AND HABITAT. Red Sea and East Africa, Madagascar and Mascarenes east to Marshall , northern Line and Tuamotu islands, north to southern Japan , south to Kimberleys (Western Australia ), Lord Howe, Norfolk and Austral islands. Found on coral and rocky reefs, including areas with freshwater runoff, 0– 30 m . Transitional water and marine. Coradion altivelis McCulloch, 1916 —Highfin coralfish STATUS AT MADANG. New record from Madang, based on specimens photographed at St. CTR 13, CTR 20. DISTRIBUTION AND HABITAT. Andaman Sea east to Philippines and Vanuatu , north to Ryukyu Islands, south to New Caledonia . Found in inshore and lagoon coral reefs, 3– 30 m . Marine. Coradion chrysozonus ( Cuvier [ex Kuhl & Hasselt] in Cuvier & Valenciennes, 1831) —Orange-banded coralfish STATUS AT MADANG. New record from Madang, based on specimens photographed at St. CTR 16, CTR 34. Andaman Sea east to Philippines and New Guinea , north to Ryukyu and Ogasawara islands, south to Solomon Islands . Found in coastal reefs, 3– 60 m . Marine. FIGURE 11 . Chaetodon trifascialis (Chaetodontidae) , Madang Lagoon, St. CTR 9, M. Hamel, 28 Oct. 2012. Coradion melanopus ( Cuvier in Cuvier & Valenciennes, 1831) —Two-eyed coralfish STATUS AT MADANG. New record from Madang, based on specimens photographed by A. Berberian and P. Laboute (St. PR 86, PR 105). DISTRIBUTION AND HABITAT. Southern Indonesia east to Philippines and New Guinea , north to Ryukyu Islands, south to Solomon Islands . Found in pairs, in coastal and seaward coral reefs, often near sponges, 10– 30 m . Marine. Forcipiger flavissimus Jordan & McGregor in Jordan & Evermann, 1898 —Forcepsfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 4, CTR 13, CTR 16, CTR 16B, CTR 19, CTR 20, CTR 22, CTR 25, CTR 28, CTR 32. CAS material (CAS 65199). DISTRIBUTION AND HABITAT. Red Sea and East Africa, Madagascar and Mascarenes east to tropical eastern Pacific, north to southern Japan , south to Western Australia , northern New Zealand , and Rapa. Found in lagoon and exposed seaward coral reefs, occasionally also on rocky reefs, 1– 114 m . Marine. Forcipiger longirostris (Broussonet, 1782) —Longnose butterflyfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 9, CTR 13, CTR 16, CTR 20, CTR 22, CTR 28, CTR 31, CTR 32, CTR 33, CTR 34. DISTRIBUTION AND HABITAT. East Africa, Comores and western Mascarenes east to Hawaiian Islands and Pitcairn Group, north to Ogasawara Islands, south to northern Australia , New Caledonia and Tonga . Found on seaward coral reefs, 3– 70 m . Marine. Hemitaurichthys polylepis (Bleeker, 1857) —Pyramid butterflyfish STATUS AT MADANG. New record from Madang, based on specimens photographed at St. CTR 13, CTR 16, CTR 22, CTR 31, CTR 32, PR 102. AMS material (AMS I.17290-001). DISTRIBUTION AND HABITAT. Indonesia east to Hawaiian and Marquesas islands and Pitcairn Group, north to southern Japan and Ogasawara Islands, south to northern Australia and New Caledonia . Found in large aggregations above outer slopes of seaward coral reefs, 3– 60 m . Marine. Heniochus acuminatus (Linnaeus, 1758) —Longfin bannerfish STATUS AT MADANG. New record from Madang, based on 2 specimens observed at Madang Harbour, 8 November 2012 . Also observed and photographed at St. CTR 0 5, CTR 16, CTR 16B; a specimen collected by W.- J. Chen on Madang Market. NTUM material (NTUM 10185). DISTRIBUTION AND HABITAT. East Africa, Persian Gulf, Madagascar and Mascarenes east to Wake and Tuamotu islands, north to southern Japan , south to Western Australia , Lord Howe, Tonga and Austral islands. Found in deep, protected lagoons and channels, and seaward coral or rocky reef slopes, 2– 178 m . Marine. Heniochus chrysostomus Cuvier [ex Parkinson] in Cuvier & Valenciennes, 1831 —Pennant bannerfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 11, CTR 12, CTR 16, CTR 20, CTR 23, CTR 32, PR 228. CAS material (CAS 65200). DISTRIBUTION AND HABITAT. Christmas and Cocos-Keeling islands (eastern Indian Ocean) east to Line Islands and Pitcairn Group, north to Ryukyu Islands, south to northern Australia and New Caledonia . Solitary or in pairs, in coastal, lagoon and seaward coral reefs, 1– 45 m . Marine. Heniochus monoceros Cuvier in Cuvier & Valenciennes, 1831 —Masked bannerfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 28, CTR 31. DISTRIBUTION AND HABITAT. East Africa, Madagascar and Mascarenes east to Marshall Islands and Henderson Island ( Pitcairn Group), north to southern Japan , south to Western Australia , Norfolk and Gambier islands. Found in lagoon and seaward coral or rocky reefs with rich coral growth, 0– 30 m . Marine. Heniochus singularius Smith & Radcliffe, 1911 —Singular bannerfish STATUS AT MADANG. First record from Madang by Kailola (1987b: 339) . Specimens observed and photographed at St. CTR 16B, CTR 30. DISTRIBUTION AND HABITAT. Maldives and Chagos Archipelago east to Caroline Islands and Samoa , north to southern Japan , south to northern Australia and New Caledonia . Solitary or in pairs, in lagoons and on seaward coral reefs, 2– 250 m . Marine. Heniochus varius (Cuvier, 1829) —Humphead bannerfish STATUS AT MADANG. New record from Madang, based on specimens observed and photographed at St. CTR 0 1, CTR 0 3, CTR 0 4, CTR 0 6, CTR 0 7, CTR 0 8, CTR 0 9, CTR 10, CTR 11, CTR 12, CTR 13, CTR 14, CTR 15, CTR 16, CTR 20, CTR 21, CTR 22, CTR 24, CTR 25, CTR 27, CTR 28, CTR 30, CTR 31, CTR 32, CTR 33, CTR 35, PR 101; a specimen collected by W.-J. Chen in Madang Bay. AMS, CAS and NTUM material (AMS I.17088- 0 60, I.17290-002; CAS 65102, 65202, 65203; NTUM 10239). DISTRIBUTION AND HABITAT. Indonesia east to Caroline and Society islands, north to southern Japan , south to northern Australia and New Caledonia . Solitary, in pairs or in groups, in coral-rich areas of lagoon coral reefs and slopes of seaward coral reefs, 2– 30 m . Marine.