Tripogon nicorae var. aristulata (Poaceae), a new variety from Peru
Author
Gutiérrez, Harol
0000-0002-0601-6121
Dirección General de Diversidad Biológica, Ministerio del Ambiente, Lima, Perú. & Universidad Científica del Sur, Lima, Perú. & gutierrezpe 5 @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0601 - 6121
gutierrezpe5@hotmail.com
Author
Castañeda, Roxana
0000-0003-3023-4740
Universidad Científica del Sur, Lima, Perú. & rcastanedas @ cientifica. edu. pe; https: // orcid. org / 0000 - 0003 - 3023 - 4740
rcastanedas@cientifica.edu.pe
Author
Gonzales, Teresa
0000-0003-2945-3874
Facultad de Ciencias de Ingeniería, Universidad Nacional de Huancavelica, Huancavelica, Perú & tgh _ quira @ hotmail. com; https: // orcid. org / 0000 - 0003 - 2945 - 3874
tgh_quira@hotmail.com
Author
Mendez, Alejandrina Sotelo-
0000-0002-6992-4821
Facultad de Zootecnia, Universidad Nacional Agraria La Molina, Lima, Perú. & asotelo @ lamolina. edu. pe; https: // orcid. org / 0000 - 0002 - 6992 - 4821
asotelo@lamolina.edu.pe
Author
Peterson, Paul M.
0000-0001-9405-5528
Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, D. C., 20013 - 7012, U. S. A. & peterson @ si. edu; https: // orcid. org / 0000 - 0001 - 9405 - 5528
peterson@si.edu
text
Phytotaxa
2021
2021-10-15
523
1
110
115
journal article
3998
10.11646/phytotaxa.523.1.7
fd46e672-7c7d-4579-a7a9-0209e50fc3c6
1179-3163
5572043
Tripogon nicorae
var.
aristulata
Gut. Peralta & P.M. Peterson
,
var. nov.
(
Figs. 1−3
).
Type:
—
PERU
.
Huancavelica
: Huaytara, San Isidro, rocky slopes with moderate to high slope, with seasonal vegetation, share habitat with populations of
Corryocactus brevistylus
(K. Schum. ex
Vaupel 1913: 17
)
Britton & Rose (1920: 66)
and
Colletia spinosissima
Gmelin (1791: 408)
,
13°59’50.28”S
,
75°15’24.06”W
,
3323 m
,
5 April 2018
,
H. Gutiérrez 169
(
holotype
MOL!, isotype MOL!).
Diagnosis:
—
Tripogon nicorae var. aristulata
differs from
T. nicorae var. nicorae
in having narrow paleas
0.4−0.5 mm
wide that are apically bilobed with two excurrent veins forming mucros
0.3–0.5 mm
long.
FIGURE 2.
Tripogon nicorae
var.
aristulata
in its natural habitat in Department Huancavelica, Peru. Photo by H. Gutiérrez (5 April 2018).
FIGURE 3.
Geographic distribution of
Tripogon nicorae
var.
aristulata
and
T. nicorae
var. nicorae
.
Caespitose perennials with fibrous remains of old sheaths. Culms (2.2–)
3–5 cm
tall. Sheaths striate, glabrous. Ligules
0.5–0.8 mm
long, membranous, densely pilose at margins, the hairs up to
0.5 mm
long. Leaf blades 15–40 ×
0.3–0.5 mm
, filiform, conduplicate, abaxially glabrous, with white and twisted long hairs on adaxial surface and two lateral tufts of long hairs near the ligule, margins scaberulous. Terminal raceme
0.8–2.4 cm
long, straight, exserted, dark olivaceous to purplish. Rachis
0.5–0.6 mm
wide, margins scaberulous. Spikelets 4–8 ×
1.2–1.5 mm
, 4–5-flowered, subsessile, alternate on the rachis, the distal ones crowded and the basal ones distant from each other. Glumes 1-veined, lanceolate, glabrous. Lower glume
2–2.5 mm
long, apex mucronate, keels scabrous. Upper glume
2.6–3 mm
long, apex obtuse. Florets imbricate. Lemmas 2.3–2.5 ×
0.6–0.7 mm
(not including awns), ovate, broadened at the middle third and narrow towards the apex, 3-veined, glabrous, apex bilobed, midvein prolonged in a straight awn, the awn
1.2–2.9 mm
long, lateral veins excurrent and mucronate, the mucros
0.4–0.6 mm
long; callus pilose, the hairs
0.5–1 mm
long; rachilla internodes
0.6–0.8 mm
long, glabrous, scar joint narrowly elliptic. Paleas 2
–
2.5 ×
0.4
–
0.5 mm
, apex bilobed with two excurrent veins forming mucros
0.3
–
0.5 mm
long. Lodicules 2, ca.
0.3 mm
long, apex truncate. Stamens 3, anthers
0.4–0.5 mm
long, purplish. Stigmas 2, plumose, purplish. Caryopsis 1–1.5 ×
0.2–0.3 mm
, subtriangular in transverse section, without ventral furrows, endosperm dry, hilum ellipsoid, embryo 1/3 the length of the caryopsis.
Ecology and distribution:
—
This
variety has been collected at high elevations in the mountains of
Peru
in the
Department
of
Huancavelica
and
Moquegua
(
Fig. 3
), on moderate to high rocky slopes, at
2472–3600 m
, with seasonal vegetation
.
Etymology:
—The epithet
aristulata
refers to the presence of two mucronate excurrent veins of the palea (
Fig. 1F
).
Taxonomical notes:
—
Tripogon nicorae
var.
aristulata
can be separated from
T. nicorae
var.
nicorae
in having apically bilobed paleas with two excurrent veins that form mucros
0.3–0.5 mm
long (
Fig. 1F
; vs. apex 2–toothed in
T. nicorae
var.
nicorae
), lemmas with lateral excurrent veins with mucros
0.4–0.6 mm
long (
Fig. 1D–E
; vs.
0.1–0.4 mm
long), and keels of the lower glume scabrous (vs. keels smooth). The typical variety of this species occurs in
Argentina
,
Bolivia
,
Chile
,
Ecuador
, and
Peru
(
Rúgolo & Vega 2004
), whereas the new variety is only known from
Peru
(
Fig. 3
).
Additional specimens examined:
—
PERU
.
Moquegua
:
Mariscal Nieto, S.C
. Calacoa, road to
Sijuaya
, rocky slope, xerophytic scrub,
16°40’16”S
,
70°44’50”W
,
2472 m
,
19 March 2019
,
D. Montesinos
7418
(MOL!)
.