Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue Author Pall-Gergely, Barna Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary https://orcid.org/0000-0002-6167-7221 pallgergely2@gmail.com Author Sajan, Sheikh Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India & Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India https://orcid.org/0000-0002-2785-6824 Author Tripathy, Basudev Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India Author Meng, Kaibaryer National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China Author Asami, Takahiro Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan https://orcid.org/0000-0001-5706-0272 Author Ablett, Jonathan D. Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom text ZooKeys 2020 981 1 220 http://dx.doi.org/10.3897/zookeys.981.53583 journal article http://dx.doi.org/10.3897/zookeys.981.53583 1313-2970-981-1 5194AAC86B8A473F8A41470A60182A0B 7C44C797C4125A71BAE032A55E6FA5DC Metalycaeus rathouisianus (Heude, 1882) Figs 34 , 35 Alycaeus rathouisianus Heude, 1882: 7, pl. 12, figs 12, 12a. Alycaeus rathouisianus - Moellendorff 1882 : 345. Alycoeus [sic] neglectus Heude, 1885: 96, pl. 24, fig. 4. syn. nov. Alycaeus (Orthalycaeus) rathouisianus - Moellendorff 1886 : 169. Alycaeus (Orthalycaeus) neglectus - Moellendorff 1886 : 170. Alycaeus (Chamalycaeus) neglectus - Kobelt and Moellendorff 1897 : 149; Kobelt 1902 : 360. Alycaeus (Chamalycaeus) rathouisianus - Kobelt and Moellendorff 1897 : 149; Kobelt 1902 : 362. Alycaeus dolomiticus Heude, 1890: 130, pl. 38, fig. 1. Chamalycaeus rathouisianus - Yen 1939 : 30, pl. 2, fig. 36. Chamalycaeus (Chamalycaeus) rathouisianus - Zilch 1957 : 145. Metalycaeus rathouisianus - Pall-Gergely et al. 2017 : 98-103, figs 55A-C, 63A, B, 65A, B. Dicharax (?) neglectus - Pall-Gergely et al. 2017 : 105-107, fig. 69K. Type locality. "E collibus juxta civitatem Song-kiang (松江), provinciae Kiang-sou, ad montes districtus Tong-lieou, sed non ubique" ( Alycaeus rathouisianus ); " ad rupestres colles civitatis Kien-te , provinciae Ngan-houe legeram " ( Alycaeus neglectus ); "Ad calcarios lapides juxta civitatem Tchen-kiang ( Kiang-sou )". Material examined. China, Tsing-p'on , MCZ 167143 (23 syntypes of A. rathouisianus ); Sung-kiang: China, coll. Moellendorff ex coll. Heude, SMF 39243 (5 syntypes of A. rathouisianus ); same data, 39303 (3 syntypes of A. rathouisianus ); China, Tchein-kiang (Kiang-sou), MCZ 167209 (29 syntypes of A. dolomiticus ); HMT-221a (labelled as lectotype of Alycaeus dolomiticus , Fig. 34 ) deposited in IZCAS: locality information were deleted; HMT-221 (labelled as paralectotype of Alycaeus dolomiticus ), same data with HMT-221; HMT-220a (1 syntype, labelled as lectotype of Alycaeus neglectus , Fig. 35 ), deposited in IZCAS: Tong-lieu (Dong-Liu town, Dong-Zhi County, Chi-Zhou City, An-Hui Province, China); Same data, HMT-220 (1 syntype, labelled as paralectotype of Alycaeus neglectus ). Remarks. Alycaeus dolomiticus Heude, 1890 is a synonym of Metalycaeus rathouisianus (see Pall-Gergely et al. 2017 ). According to Johnson (1973) , paratypes of A. neglectus are present in the MCZ (inv. number: 167222) and the USNM (inventory number: 472336). In the revision of Chinese Alycaeidae ( Pall-Gergely et al 2017 ) we examined the former sample, but the locality ("China, Tong-lieou" in Anhui Province) did not match the type locality given in the original description ( "Kien-te" in Zhejiang Province). Consequently, we did not consider them as type specimens; moreover, on a hand-written label of the MCZ 167222 sample the following text was written: "not types, only holotype in Heude's coll.". We have now been able to examine the specimens in the Beijing museum, and found that the specimens labelled as types also have the same type locality ( "Tong-lieu" ). The Beijing specimens were also labelled as lectotype (HMT-221a), and paralectotypes (HMT-221). It appears that although the locality on the type specimen labels and that in the original description do not match, the shells labelled as types are indeed types of A. neglectus . Since we did not find traces of the lectotype designation, we consider all type specimens of A. neglectus syntypes. Since we did not consider the specimens in the MCZ types in Pall-Gergely et al. (2017) , we had to rely on the original description regarding the identity of A. neglectus , and based on the low protoconch in the figure of the original description, we tentatively classified it in Dicharax . However, this opinion must now be revised: the shells labelled as types in the Beijing Museum and in the MCZ are identical, and they are also identical to typical Alycaeus rathouisianus shells. Thus, A. neglectus is considered a synonym of A. rathouisianus . Figure 34. Metalycaeus rathouisianus (Heude, 1882), syntype: labelled as lectotype of Alycaeus dolomiticus (HMT-221a). Photographs: Kaibaryer Meng. Figure 35. Alycaeus neglectus Heude, 1885, syntype (HMT-220a; synonym of Metalycaeus rathouisianus (Heude, 1882)). Photographs: Kaibaryer Meng.