Species diversity in the Monodelphis brevicaudata complex (Didelphimorphia: Didelphidae) inferred from molecular and morphological data, with the description of a new species
Author
Pavan, Silvia Eliza
Author
Rossi, Rogerio Vieira
Author
Schneider, Horacio
text
Zoological Journal of the Linnean Society
2012
2012-04-18
165
1
190
223
http://dx.doi.org/10.1111/j.1096-3642.2011.00791.x
journal article
10.1111/j.1096-3642.2011.00791.x
0024-4082
5407298
MONODELPHIS MARAXINA
THOMAS, 1923
Monodelphis maraxina
Thomas, 1923
was described for Marajó Island, in eastern state of
Pará
.
Thomas (1923)
suggested that
M. maraxina
is closely related to
M. domestica
. Subsequent authors agreed with him, some of which even regarded
M. maraxina
as a synonym of
M. domestica
(e.g.
Cabrera, 1958
;
Emmons & Feer, 1990
,
1997
). However,
Pine (1980)
suggested that a relationship of
M. maraxina
with members of the
M. brevicaudata
group would be more reasonable on zoogeographical grounds. He also noted that the relative hairlessness of the tail and the buffy coloration of body sides are conditions shared with specimens of
M. brevicaudata
from the east bank of the Tapajós river (=
M. glirina
). Recently, based on
Pine’s (1980)
view,
Solari (2010)
included
M. maraxina
in the
M. brevicaudata
group.
Figure 8.
Bivariate scatterplot from the canonical discriminant analysis. Wilks’ lambda = 0.093,
F
= 12.830, d.f. = 1.696;
P
<0.001.
Our morphological analysis of the
holotype
of
M. maraxina
(BMNH 12.5.11.13) through photographs led us to conclude that this form undoubtedly belongs to the
M. brevicaudata
complex. The colour pattern observed in specimens of
M. glirina
from the east bank of the Xingu river shows a striking concordance with the original description of
M. maraxina
, and our analysis confirmed that these two forms have highly similar skin and skulls. Similarities between the faunas of the Marajó Island and the adjacent area at the eastern margin of the Xingu river are expected, as indicated by other vertebrate groups (e.g.
Nascimento
et al
., 1991
;
Avila-Pires, 1995
;
Fernandes, Cardoso da Silva & Silva Jr, 1995
;
Ron, 2000
). Furthermore, our recent field work at Marajó uncovered specimens of another species of the
M. brevicaudata
complex that is also present on the east bank of the Xingu river (
M. touan
), confirming that these areas are able to share species of short-tailed opossum.
Given the substantial morphological similarity between the
holotype
of
M. maraxina
and specimens of
M. glirina
from east of the Xingu river, and the evidence that these areas are able to share species of short-tailed opossum, we argue that
M. maraxina
may represent a geographical variant of
M. glirina
. However, as we have not performed molecular and direct morphological analyses of specimens regarded as
M. maraxina
, and because our general analyses are not conclusive with respect to
M. glirina
, which may represent a complex of species rather than an unique species, more accurate analyses are needed to answer this question. At this time, we maintain
M. maraxina
as a valid species included in the
M. brevicaudata
complex.