Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)
Author
Ruud, A. Bank
Author
Henk P. M. G Menkhorst
Author
Eike Neubert
text
Basteria
2016
80
1
5
30
journal article
10.5281/zenodo.439745
e7d164f6-b22c-4120-b1f6-6464cfc6de0b
00056219
439745
Familia
Vitrinidae Fitzinger, 1833
From the European mainland, Turkey, the Caucasus and northern Africa (Libya and Algeria) several vitrinid genera have been reported, encompassing about 30 species. These are the genera
Eucobresia
H.B. Baker, 1929
,
Hessemilimax
Schileyko, 1986,
Oligolimax
P. Fischer, 1878
,
Phenacolimax
Stabile, 1859
,
Sardovitrina
Manganelli & Giusti, 2005
,
Semilimacella
Soós, 1917
,
Semilimax
Gray, 1847
,
Vitrina
Draparnaud, 1801
, and
Vitrinobrachium
Künkel, 1929
. None of the species from these genera have a shell with a maximum diameter above 10 mm. The situation is different with respect to Macaronesia (Madeira, Canary Islands, Azores). It is home for 35 endemic species classified into the genera
Azorivitrina
Giusti, Fiorentino, Benocci & Manganelli, 2011,
Canarivitrina
Valido & Alonso, 2000
,
Guerrina
Odhner, 1955
,
Insulivitrina
P. Hesse, 1923
,
Madeirovitrina
Groh & Hemmen, 1986
, and
Plutonia
Stabile, 1864
. Shells of a number of vitrinids of Macaronesia are relatively large, and reach sizes above 11 mm:
Insulivitrina
and
Madeirovitrina
.
Calidivitrina
Pilsbry, 1919, is a genus endemic for the African continent. Representatives of this genus are known from Kenya, Uganda, Tanzania and the eastern border of Congo-Kinshasa. These are:
bambuseti
(Thiele, 1911),
cagnii
(Pollonera, 1906),
chyuluensis
(
Verdcourt, 2005
),
ericinellae
(d’Ailly, 1910),
ibandensis
(Pollonera, 1907),
kiboschoensis
(d’Ailly, 1910),
lactae
Connolly, 1925,
lobeliaecola
(Dautzenberg, 1908),
nigrocincta
(E. von Martens, 1887),
oleosa
(E. von Martens, 1895) [= type species],
ugandensis
(Thiele, 1911),
tenuissima
(Thiele, 1911),
variopunctata
(Connolly, 1931), and
viridisplendens
(d’Ailly, 1910). The maximum diameter of the shell of most
Calidivitrina
species is in the range between 8 and 11 mm; the largest two species are
bambuseti
(12 mm) and
chyuluensis
(13 mm).
Figs 16-20.
Multidentula
and
Imparietula
taxa from Turkey.
16
,
Multidentula squalina squalina
(L. Pfeiffer, 1848)
, NMBE 548313 (H = 7.8 mm), Turkey, Vil. Karabük, eastern part Eskipazar, leg. Hausdorf, 27.ix.1987.
17
,
Multidentula squalina eudoxina
(Nägele, 1894)
, paralectotype SMF 14195a coll. Kobelt ex Nägele (H = 6.3 mm), Turkey, Vil. Tokat, Tokat.
18
,
Multidentula squalina acutior
(Lindholm, 1922)
, NMBE 544678 (H = 5.7 mm), Turkey, Vil. Artvin, N. Kınalıçam (= NNW. Cağliyan).
19
,
Multidentula reducta
spec. nov.
, holotype NMBE 544681 (H = 6.2 mm), Turkey, Vil. Sivas, Zara, 16 km towards Imranlı.
20
,
Imparietula inflexa
spec. nov.
, holotype NMBE 544683 (H = 12,4 mm), Turkey, Vil. Giresun, 5 km N. Tamdere (= S. Yavuzkemal). All photographs Bochud & Neubert, × 8.
The genus
Arabivitrina
Thiele, 1931 is distributed in Yemen and in the southwestern part of Saudi Arabia, namely
A. arabica
(Thiele, 1910) [= type species] and
A
.
jansseni
Neubert, 1998
, as well as in Abyssinia (Ethiopia). The two taxa from Saudi Arabia / Yemen have been reviewed by
Neubert (1998)
; the shell reaches a maximum diameter of about 21 and 18 mm, respectively. From Ethiopia a large number of taxa have been described that might be assigned to
Arabivitrina
. These are
abyssinica
(L. Peiffer, 1848),
antonellii
(Pollonera, 1888),
bianchii
(Pollonera, 1888),
caillaudi
(Morelet, 1872),
chiauzzii
(Bacci, 1940),
conquisita
(Jickeli, 1882),
darnaudi
(L. Pfeiffer, 1856),
demissa
(Pollonera, 1898),
devexa
(Jickeli, 1873),
giuliettii
(Pollonera, 1888),
grossepunctata
(Connolly, 1928),
helicoidea
(Jickeli, 1873),
herbini
(Bourguignat, 1883),
humilis
(Pollonera, 1898),
isseli
(Morelet, 1872),
jamjamensis
(Kobelt, 1905),
jickelii
(Jickeli, 1874),
licatae
(Pollonera, 1888),
martensii
(Jickeli, 1873),
martinii
(Pollonera, 1888),
milneedwardsiana
(Bourguignat, 1883),
modesta
(Pollonera, 1898),
naticoides
(Pollonera, 1888),
neumanni
Thiele, 1933,
olivacea
Thiele, 1933,
perobliqua
(Innes, 1884),
planulata
(Jickeli, 1873),
raffrayi
(Bourguignat, 1883),
ragazzii
(Pollonera, 1888),
riepiana
(Jickeli, 1882), and
semirugata
(Jickeli, 1873). However, the validity of these taxa is in most cases quite unclear: it is reasonable to assume that, after revision, several names turn out to be synonyms. Furthermore, it is even possible that some names do not belong to
Arabivitrina
, but to members of the superfamily Helicarionoidea. Most of the mentioned taxa have relatively large dimensions, i.e. the maximum diameter of the shell is in the range between 11 and 21 mm. Two species are exceptionally large, namely
licatae
and
ragazzii
, with reported sizes of 23 and 30 mm, respectively. For an introduction into the chaotic state of the African
Vitrinidae
we refer to Forcart (1956, 1978),
Hubendick (1953)
and
Verdcourt (2005)
.
Three vitrinid species have been reported from Madagascar:
madagascariensis
E.A. Smith, 1882,
marojeziana
Fischer-Piette, C. Blanc,
F. Blanc & Salvat, 1994
, and
josephinae
Emberton & Griffiths, 2009
. However, the original figures of
marojeziana
and
josephinae
show an animal with a caudal horn; these taxa should therefore be assigned to the superfamily Helicarionoidea, as will undoubtedly be the case for
madagascariensis
as well.
We stumbled upon a large Turkish vitrinid species collected at the Nemrud Dağ, which we cannot assign to one of the above mentioned genera. A most likely closely related -but almost forgotten- species lives at several localities in Lebanon, currently known as
Vitrina libanica
Germain, 1911
.