Sitticine jumping spiders: phylogeny, classification, and chromosomes (Araneae, Salticidae, Sitticini) Author Maddison, Wayne P. Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada https://orcid.org/0000-0003-4953-4575 wmaddisn@mail.ubc.ca Author Maddison, David R. Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA https://orcid.org/0000-0002-7152-3824 Author Derkarabetian, Shahan Department of Biology, San Diego State University, San Diego, CA 92182, USA & Department of Organismic and Evolutionary Biology, Harvard University, Cambridge MA 02138, USA https://orcid.org/0000-0002-9163-9277 Author Hedin, Marshal Department of Biology, San Diego State University, San Diego, CA 92182, USA text ZooKeys 2020 925 1 54 http://dx.doi.org/10.3897/zookeys.925.39691 journal article http://dx.doi.org/10.3897/zookeys.925.39691 1313-2970-925-1 BB966609087849A1B13C138C2495E6B7 DCC1953545B9557EA73102AE6D25711F Tomis manabita W. Maddison sp. nov. Figures 120-128 Type material. Male holotype, 10 male and 8 female paratypes from Ecuador: Manabi : Puerto Rico, Cabanas Alandaluz 5 May 1994 W. Maddison WPM#94-031. The holotype (specimen UBC-SEM AR00217) pertains to the Museum of Zoology, Pontificia Universidad Católica, Quito, Ecuador (QCAZ), but is currently held in the Spencer Entomological Collection at the Beaty Biodiversity Museum, University of British Columbia (UBC-SEM). Etymology. Based on the type locality; the form is the adjective in Spanish (masculine or feminine). A species on the beaches of coastal Ecuador, resembling Attulus in habitus. It was used in the molecular phylogenetic study of Maddison and Hedin (2003) under the name " Sitticus sp." (voucher S220) from Manabi , Ecuador. Diagnosis. Palp closely resembles that of Tomis pavidus , from which it differs in the smaller tibia and longer RTA. Description. Male (holotype). Carapace length 1.58; abdomen length 1.51. Carapace (Figs 124 , 127 ) medium brown with recumbent brown setae except for thin medial longitudinal band of white setae on thorax, two spots of pale setae in ocular area, and a marginal band of white setae that is broad on the thorax but narrows to the anterior, disappearing before the clypeus is reached. Clypeus brown, with a few brownish setae. Chelicerae vertical, with a few pale setae near the clypeus. Retromarginal cheliceral teeth lacking. Palp clothed with white setae dorsally, but prolaterally with darker integument and setae from tip of femur to cymbium; cymbium mostly dark brown. Embolus (Fig. 120 ) arises broadly, more centrally beneath bulb (and less peripherally) than is typical, then narrows abruptly at ca. 9 o'clock . RTA extremely long and thin, parallel to axis of the palp. Legs honey-coloured, with notably darker annulae at the tarsus-metatarsus joints, and black stripe on prolateral-ventral face of first patella and tibia. Abdomen (Figs 124 , 127 ) brown with two lateral and one median longitudinal bands of paler setae, the medial band less distinct, wavy, and accompanied by a lateral extension that forms a cross. Female (paratype # UBC-SEM AR00218). Carapace length 1.76; abdomen length 2.22. Carapace (Figs 126 , 128 ) brown, covered unevenly with recumbent cream-coloured setae. Clypeus with white setae, longest at midline where they overhang the chelicerae. Chelicerae brown, with a few setae near clypeus. Palps and legs honey coloured, with weak annulae. Abdomen brown, marked as in male except bands are less distinct (Figs 126 , 128 ). Epigynum with an anterior atrium from which short copulatory ducts lead diagonally to the spermathecae (Figs 122 , 123 ). Additional material. 15 males and 7 females from Ecuador: Manabi : Machalilla National Park, Salaite, between HWY and coast 6 May 1994 W. Maddison WPM#94-032 (4 males, 2 females); Ecuador: Manabi : Machalilla National Park, Salaite, 1 km inland along trail from HWY. 6 May 1994 W. Maddison WPM#94-033 (3 males); Ecuador: Manabi : Machalilla National Park, trail between Agua Blanca & San Sebastien 50-400 m dry forest 7 May 1994 W. Maddison WPM#94-034 (1 male); Ecuador: Manabi : Crucita. 30 August 1988 W. Maddison WPM#88-040 (2 males 4 females); Ecuador: Del Oro: Jambeli 13 August 1989 W. Maddison WPM#89-040 (3 males); Ecuador: Manabi : Puerto Lopez. 1.5497 , -80.8104 ; 5 m el. 1-5 August 2004 W. Maddison. WPM#04-067 (2 males 1 female).