Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary pallgergely2@gmail.com Author Hunyadi, Andras Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary Author Vermeulen, Jaap J. JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands Author Grego, Jozef Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia Author Sutcharit, Chirasak Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Reischuetz, Alexander Puechhaimgasse 52, A- 3580 Horn, Austria Author Dumrongrojwattana, Pongrat Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand Author Botta-Dukat, Zoltan Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary Author Oerstan, Aydin 12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA Author Fekete, Judit University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary Author Jochum, Adrienne https://orcid.org/0000-0002-6624-6412 Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany text ZooKeys 2023 2023-02-13 1147 1 177 http://dx.doi.org/10.3897/zookeys.1147.93824 journal article http://dx.doi.org/10.3897/zookeys.1147.93824 1313-2970-1147-1 9BB9881B0076473D8E53155D37CA1F50 FF2B6B317B505F9EA0E1000BDCD16CE7 Angustopila antidomedon Pall-Gergely & Hunyadi sp. nov. Fig. 61 Type material. Holotype : Vietnam • 1 empty shell (H: 0.94 mm, D: 0.83 mm); Thanh Hoa Province, Như Thanh District, Hải Van , Hang Lo Cao Khang Chiến, vicinity of the cave (locality code: 2020/41); 19°37.08'N , 105°34.63'E ; 20 m a.s.l.; 14 Feb. 2020; A. Hunyadi leg.; HNHM 105294. Paratypes : Vietnam • 1 imaged + 8 other shells; same data as for holotype; coll. HA. Diagnosis. A medium-sized, conical Angustopila species with a narrow umbilicus, a parietal tooth and a blunt but strong palatal tooth situated at the lower part of the palatal side. Description. Shell of normal size for the genus, higher than wide; off-white, conical; body whorl widest from standard apertural view; protoconch consists of 1.25 whorls, with weak spiral striation preceding the first teleoconch whorl; teleoconch with rather strong, irregularly spaced radial growth lines and equally strong, equidistantly and sparsely-arranged spiral striae (ca. 12-19 on body whorl from standard apertural view); whorls 4.5 or slightly less, rounded or very slightly depressed-rounded in form; aperture oblique to shell axis from lateral view; umbilicus narrow; aperture subovoid or reniform with very wide, not distinctly separated sinulus; peristome expanded, not reflected; parietal callus slightly protruding, somewhat detached from penultimate whorl; parietal tooth straight, strong, normally high and long, situated inside shell at some distance from peristome edge; lower palatal tooth, blunt but strong; approximately half of the available shells has a tiny subcolumellar tooth. Measurements (in mm). H = 0.86-0.94; D = 0.82-0.88, H/D*100 = 103.6-113.3 ( n = 4), RUD = 24.2-25.0 ( n = 2). Differential diagnosis. Angustopila pustulata sp. nov. is generally larger (wider), although the size ranges slightly overlap (D of A. antidomedon sp. nov.: 0.82-0.88 mm, D of A. pustulata sp. nov.: 0.87-0.93 mm), it has a lower spire (Fig. 77A is a rare example, nearly all shells have a low spire, similar to the holotype), and its palatal tooth is much lower (weaker). Moreover, some shells of A. antidomedon sp. nov. possess a small columellar tooth. Angustopila occidentalis sp. nov. is larger, has a lower spire and a comparatively larger aperture with usually weaker palatal tooth. Angustopila uvula sp. nov. has a concave-conical shell and a stronger parietal lamella. See also under A. bathyodon sp. nov. and A. gracilis sp. nov. Figure 61. Angustopila antidomedon Pall-Gergely & Hunyadi, sp. nov. A, C-F, I, J holotype (HNHM 105293) B, H paratype (specimen 2). Apertural ( A, B ), lateral ( C ), apical ( D ) and ventral ( E ) sides of the shell; sculpture on the protoconch ( F ), aperture ( G, H ); ventral ( I ) and frontal ( J ) surface of the body whorl. Etymology. From the Greek ἀντιδομή (= opposed) and ὀδούς (= tooth) referring to the position of the palatal tooth in relation to the parietal tooth. Distribution. This species is known from the type locality only (Fig. 27 ).