Review of the genus Hypostomus Lacépède, 1803 from rio Ribeira de Iguape basin, with description of a new species (Pisces, Siluriformes, Loricariidae).
Author
Osvaldo T. Oyakawa
Author
Alberto Akama
Author
Angela M. Zanata
text
Zootaxa
2005
921
1
27
http://www.zoobank.org/urn:lsid:zoobank.org:pub:913A8172-1A2E-4784-96DB-50BACBEC7C25
journal article
z00921p001
913A8172-1A2E-4784-96DB-50BACBEC7C25
[[ Genus
Hypostomus Lacépède
]]
Hypostomus
, the type genus of the subfamily Hypostominae, constitutes an assemblage of bottom-dwelling loricariid fishes widely distributed throughout South America, occurring in a variety of freshwater ecosystems such as small mountain streams and large lowland river areas. Maximum diversity in number of species of
Hypostomus
occurs in rivers of the
Parana-Paraguay
system (Weber, 2003). As presently defined the genus consists of more than 130 nominal species (107 species considered valid by Weber, 2003 and expanded by Armbruster, 2004).
Species level taxonomy of
Hypostomus
is still poorly known as well as the phylogenetic relationships within and of the genus.
Hypostomus
was established by
Lacepede
(1803), who included in the genus only a single species,
H. guacari
. Old descriptions, similar to
Lacepede
’s, and available for various subsequent species added to the genus, are very often scanty, with incomplete or poor characterization of species. This fact, together with a number of
Hypostomus
species yet to be described, makes the recognition of most taxa of the genus difficult. Hypothesis of phylogenetic relationships of
Hypostomus
based on molecular data (Montoya-Burgos et al., 1997, 2002) revealed that the genus does not form a natural group. According to these authors, some species currently assigned to
Hypostomus
should be removed from the genus, while the genus
Cochliodon
should be treated as synonym of
Hypostomus
, an action effectuated by Weber & Montoya-Burgos (2002: 265). Morphological based phylogenetic studies presented by Armbruster (2004) reinforce the conclusions above and place
Aphanotorulus
and
Isorineloricaria
in the synonymy of
Hypostomus
. In spite of the cited studies, more detailed phylogenetic analysis of the relationships within
Hypostomus
awaits for future studies.
The elevated number of species currently included in
Hypostomus
, allied to the broad variability in morphology and color pattern of those taxa, poses difficulties for its entire revision. Various authors acquainted with these problems provided regional revisions of the genus, as exemplified by Boeseman (1968) with a study of the Surinam representatives of the genus, Weber (1985, 1986, 1987) that deals with
Hypostomus
from Paraguay, more recently Reis et al. (1990) reviews and describes three new species from Southern Brazil, Mazzoni et al. (1994) provides a taxonomical revision of
Hypostomus
from Lower rio
Paraiba
do Sul, State of Rio de Janeiro, Brazil, and finally Hollanda Carvalho & Weber (2004) provided a revision of
Hypostomus cochliodon group
from middle and lower Amazon system.