The pill millipedes of Vietnam: a key to genera and descriptions of five new species (Diplopoda: Glomerida: Glomeridae) Author Nguyen, Anh D. Author Sierwald, Petra Author Marek, Paul E. text Raffles Bulletin of Zoology 2019 2019-04-16 67 260 297 journal article 10.26107/RBZ-2019-0020 2345-7600 4575924 C1D8A501-317A-4803-AEF8-EA16DFBFF3B2 Hyperglomeris simplex , new species (Figs. 19–23) Material examined. Holotype : male ( FMNH INS 3716071 ) Vietnam , Vinh Phuc Province , Phuc Yen Town , Ngoc Thanh Commune , Me Linh Station for Biodiversity , 21.3850°N , 105.7119°E , regenerated forest, 10–16 September 2016 , coll. Petra Sierwald et al. Paratypes : 1 male , 1 female ( FMNH INS 3716070 ) same as for holotype ; 2 females (IEBR–Myr 605) same locality, but on the way to Tam Dao 2, natural forests, 1,100 m , 25 February 2017 , leg. Anh D. Nguyen ; 1 female (IEBR–Myr 655) same data as for IEBR–Myr 605, but 2–4 August 2017 . Etymology. Named after the simple tibial knob of the male telopods (feminine adjective). Diagnosis. The species is differentiated by the following character combination: Terga black with median line of yellow spots longitudinally, and two paramedial marbled transverse oval spots. Femoral trichostele absent, femur with mesoanteriorly oriented lamella. Tibial process a simple knob, tibia with mesoposteriorly oriented lamella. Tarsus hooked distally. The new species differs from H. nigra Golovatch, 2017 , H. depigmentata Golovatch, Geoffroy & VandenSpiegel, 2013 in colouration pattern: body entirely unpigmented in H. depigmentata , and dorsa entirely black in H. nigra vs. terga with two lateral marble yellowish brown Fig. 19. Hyperglomeris simplex new species from Tam Dao National Park, habitus. Photo by: Anh D. Nguyen, image not to scale. spots and a medioposterior, brownish yellow, triangular spot in H. simplex , new species . It can also be distinguished from H. maxima Golovatch, 1983 , H. conspicua Golovatch, 1983 , H. dirupta ( Silvestri, 1917 ) and H. lamellosa Silvestri, 1917 in telopod conformation. Hyperglomeris maxima and H. dirupta have a very short, tiny, femoral trichostele which is absent in H. simplex , new species , H. conspicua and H. lamellosa . The latter three species are distinguished by the tibial structure: H. simplex has a simple knob whereas H. conspicua has a conspicuous, serrate, long process and a small, setiferous knob; H. lamellosa has two different processes. Fig. 20. Hyperglomeris simplex new species , holotype (FMNH INS 3716071) from Vinh Phuc Prov., northern Vietnam. A, whole body, dorsal view; B, lateral view; C, ventral view; D, head, subventral view. Scale bar = 1 mm. Fig. 21. Hyperglomeris simplex new species , holotype (FMNH INS 3716071) from Vinh Phuc Prov., northern Vietnam. A, whole body, under UV light, dorsal view; B, lateral view; C, ventral view; D, head, subventral view. Scale bar = 1 mm. Fig. 22. Hyperglomeris simplex new species , holotype (FMNH INS 3716071) from Vinh Phuc Prov., northern Vietnam. A, antenna; B, leg 17; C, legs 18; D, telopods, anterior view; E, posterior view. Scale bar = 1 mm. Description. Holotype male (FMNH INS 3716071). Width of 2nd tergum 7.0 mm, body length 12 mm . Exoskeleton : Terga blackish brown, shiny and smooth. Each tergum with two lateral light-brown marbled, transverse oval spots, and a medio-posterior, light-brown triangular spot. Caudal and lateral tergal margins light-yellow (Figs. 19 – 21). Head : Ocelli 7+1, lenses convex, black contrasting against brown background of head. Tömösváry organ transverse oval, margins not concave, ca. 1.6× wider than long ( Figs. 20C , 21C ). Antennomere 6 bean-shaped ( Fig. 22A ), not clavate, longer than antennomeres 4+5 combined. Antennal tip with four large apical sensory cones (sc). Tergites : Collum with a large oval marbled spot centrally, brown background, with 5 transverse striae ( Figs. 20D , 21D ). Second tergum with a narrow hyposchism reaching caudal margin, with 13 superficial transverse striae. Anal shield broadly rounded at caudal margin ( Figs. 20C , 21C ). Legs : Leg-pair 17 strongly reduced in size, with 4 podomeres: coxa with a rounded outer lobe (col), and with a setiferous tubercle distomesally; podomere 1 with a small distomesal setiferous knob. Leg-pair 18 ( Fig. 22C ) almost normal size, with 4 podomeres with a simple syncoxial notch, and pronounced syncoxial tubercles. Telopods ( Figs. 22D, E , 23 A–D) with a trapeziform, medially notched (arrow), sparsely setose, central syncoxial lobe (syl) with two setose horns (syh) paramedially, each directed mesoventrad. Prefemur ( Fig. 22E ) somewhat broadened basally, carrying a tuberculiform trichostele distomesally (pret); lateroanterior side with dense micropapillae (visible on Fig. 23B ). Femur simple, without trichosteles, with a large, straight, tuberculiform distal process (dfp). Tibia membranous mesally, with a short, stout knob distomesally on anterior side ( Fig. 23A , tp). Tarsus apically acuminate, hooked distally. Variation. Width of second tergum 7.0– 7.2 mm (males), 9.2 mm (female); body length 12.0– 13.2 mm (males), 14.6 mm (female).