World reclassification of the Cardiophorinae (Coleoptera, Elateridae), based on phylogenetic analyses of morphological characters Author Douglas, Hume B. text ZooKeys 2017 655 1 130 http://dx.doi.org/10.3897/zookeys.655.11894 journal article http://dx.doi.org/10.3897/zookeys.655.11894 1313-2970-655-1 8C475FAB25E044CEA2FBC3B83F316D8C 8C475FAB25E044CEA2FBC3B83F316D8C Cardiophorus: subgenus Perrinellus Buysson, 1899 Figs 98-99 Cardiophorus : subgenus Perrinellus Buysson, 1899: 282. Type species: Athous argentatus Abeille de Perrin, 1894: 92. Cardiophorus : subgenus Lasiocerus Buysson, 1912: 129. Type species: Cardiophorus schusteri Buysson, 1912: 128. Diagnosis. Head. Mandibles with 2 apices; supra-antennal carina without longitudinal split next to compound eyes. Prothorax. Pronotum with lateral carina reaching less than halfway to anterior edge (in type species not distinguishable from the dorsal hind angle carina), not hidden by lateral expansion of pronotum in dorsal view; prosternum with anterior edge produced as lobe, concealing labium when head not extended. Pterothorax. Scutellum with anterior edge emarginate, anterolateral edges broadened posterior to anterolateral corners, posterior end pointed. Legs. tarsi without ventral lobes and tarsal claws with one apex per side. Also. Pronotum with hind angles not truncate dorsally; procoxal cavities open. Aedeagus with parameres approximately cylindrical pre-apically (Fig. 99). Type locality. Israel, other spp from North Africa, Ceylon, Central Asia, probably not monophyletic, 12 spp. Cardiophorus (Lasiocerus) du Buysson was not located for examination, and may not match these key characteristics. Subgenus Lasiocerus was described from Azerbaijan, and distinguished by its long antennae with dense setae.