Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from Southeast Asia (Araneae, Zodariidae) 3369
Author
Dankittipakul, Pakawin
Author
Jocqué, Rudy
Author
Singtripop, Tippawan
text
Zootaxa
2012
2012-07-04
3369
1
1
327
https://biotaxa.org/Zootaxa/article/view/zootaxa.3369.1.1
journal article
10.11646/zootaxa.3369.1.1
11755334
5252563
Mallinella multicornis
sp. nov.
(
Figs 210–211
,
216–217, 219–220
,
230
,
240
,
273–277
)
Material
examined.
Holotype
:
♂
,
MALAYSIA
,
Penang
Island
:
Penang Hill
,
150–330 m
,
2 December 1991
, leg.
P.J. Schwendinger
(
MHNG
,
PM
–4)
.
Paratype
:
1♀
, same data as holotype (
MHNG
)
.
Diagnosis.
Males of
M. multicornis
sp. nov.
are recognized by a broad embolus in combination with the TA on male the palp provided with several teeth (
Fig. 273
). Females are most likely to be confused with those of
M. linguiformis
sp. nov.
in the similar shape of the epigynal plate (
Figs 230
cf. 229) but the apex of the spermathecae is enlarged, and subspherical (
Figs 275
cf. 295).
Etymology.
The specific epithet is a combination of Latin words (adjective,
multus
= many; adjective,
cornis
= horned) and refers to the TA on the male palp.
Description.
Male
(
holotype
). Total length 10.70; prosoma 5.28 long, 4.55 wide. Eye sizes and interdistances: AME 0.32, ALE 0.24, PME 0.20, PLE 0.20, AME–AME 0.16, AME–ALE 0.30, PME–PME 0.22, PME–PLE 0.48; MOQ: 0.62 long, 0.60 anterior width, 0.54 posterior width. Leg measurements: I 11.19 (3.07, 3.44, 2.46, 2.21), II 10.94 (2.82, 3.44, 2.70, 1.96), III 10.82 (2.95, 3.32, 2.82, 1.72), IV 13.18 (3.07, 4.30, 4.05, 1.74).
FIGURES 221–226. 221–223.
Mallinella hilaris
comb. nov.
, male (NHMV).
224–225.
M. acanthoclada
sp. nov.
, holotype.
226.
M. bidenticulata
sp. nov.
, holotype.
221, 224, 226.
Male palp, ventral.
222.
Ditto, prolateral.
223, 225.
Ditto, retrolateral.
FIGURES 227–234. 227.
Mallinella belladonna
sp. nov.
, holotype.
228.
M. bidenticulata
sp. nov.
, paratype.
229.
M. linguiformis
sp. nov.
, holotype.
230.
M. multicornis
sp. nov.
, paratype.
231.
M. rostrata
sp. nov.
, paratype.
232.
M. acanthoclada
sp. nov.
, paratype.
233.
M. hilaris
comb. nov.
, female from Java.
234.
M. calicoanensis
sp. nov.
, holotype.
227–234.
Epigyne.
Coloration
(
Fig. 210
). Carapace dark reddish brown, margin dark brown. Sternum and chelicerae reddish brown. Legs yellowish. Dorsum of opisthosoma sepia. Dorsal pattern: first pair almost indistinct; second pair represented by narrow bands extending laterally; third to fifth pairs by transverse chevrons.
Palp
(
Fig. 273
). Retrolateral margin of palpal tibia well-developed, subtriangular, apex bluntly pointed. RTA digitiform, short and stout, apex rounded. TA with elongated apico-prolateral process; apical ridge sharply pointed; mesal ridge with three teeth directed anteriad. Tegular spine pronounced, relatively broad at base, terminally blunt. Embolic base originating at 270°, anterior margin excavated to accommodate broad membranous part. Embolus broad, semi-transparent, dilated subapically.
Female
(
paratype
).
Total length 12.30; prosoma 5.94 long, 4.30 wide. Eye sizes and interdistances: Eye sizes and interdistances: AME 0.30, ALE 0.24, PME 0.20, PLE 0.22, AME–AME 0.14, AME–ALE 0.28, PME–PME 0.20, PME–PLE 0.50; MOQ: 0.60 long, 0.58 anterior width, 0.56 posterior width. Leg measurements: I 10.20 (2.95, 3.07, 2.58, 1.59), II 10.73 (2.90, 3.38, 2.58, 1.86), III 10.25 (2.97, 3.27, 2.41, 1.59), IV 10.99 (3.07, 3.17, 2.95, 1.79).
Coloration
(
Fig. 211
). Carapace reddish brown. Sternum and chelicerae brown. Legs yellowish. Dorsum of opisthosoma sepia, mottled with numerous minute pale spots. Dorsal pattern: first pair represented by elongateovoid spots; second pair by thin transverse bands, almost fused medially, extending laterally; third to fifth pairs by broad chevrons.
FIGURES 235–242. 235.
Mallinella belladonna
sp. nov.
, holotype.
236.
M. bidenticulata
sp. nov.
, paratype.
237.
M. linguiformis
sp. nov.
, holotype.
238.
M. calicoanensis
sp. nov.
, holotype.
239.
M. rostrata
sp. nov.
, paratype.
240.
M. multicornis
sp. nov.
, paratype.
241.
M. hilaris
, female from Java.
242.
M. acanthoclada
sp. nov.
, paratype.
235–242.
Internal genitalia, anterior.
FIGURES 243–249. 243–246.
Mallinella hilaris
comb. nov.
, male (NHMV).
247–249.
M. hilaris
comb. nov.
, female from type locality, Cibodas.
243.
TA, ventral.
244.
Ditto, prolateral.
245.
Ditto, retrolateral.
246.
Apex of embolus, ventral.
247.
Right spermatheca, anterior.
249.
Ditto, posterior.
248.
Left spermatheca, lateral.
FIGURES 250–255. 250–253.
Mallinella rostrata
sp. nov.
, holotype.
254–255.
M. rostrata
sp. nov.
, paratype.
250.
Male palp, ventral.
251.
TA, prolateral.
252.
Ditto, retrolateral.
253.
Ditto, ventral.
254.
Left spermatheca, anterior.
255.
Ditto, posterior.
FIGURES 256–264. 256–260.
Mallinella bidenticulata
sp. nov.
, holotype.
261–264.
M. bidenticulata
sp. nov.
, paratype.
256.
Male palp, ventral.
257.
Palpal tibia, retrolateral.
258.
TA, retrolateral.
259.
Ditto, prolateral.
260.
Apex of embolus, prolateral.
261.
Internal genitalia, dorsal.
263.
Ditto, latreral.
262.
Right spermatheca, anterior.
264.
Apex of right spermatheca, posterior.
FIGURES 265–272. 265–268.
Mallinella acanthoclada
sp. nov.
, holotype.
269–272.
M. acanthoclada
sp. nov.
, paratype.
265.
Male palp, ventral.
266.
Ditto, retrolateral.
267.
TA, prolateral.
268.
Ditto, ventral.
269.
Internal genitalia, dorsal.
270.
Right spermatheca, anterior.
271.
Left spermatheca, lateral.
272.
Ditto, posterior.
Genitalia
(
Figs 230
,
240
,
274–277
). Epigynal plate rectangular, posterior margin slightly excavated in center, lobular lateral margins strongly pronounced. Spermathecae with cylindrical proximal part, abruptly bent posteriorly, distal part rounded.
Natural history.
Mallinella multicornis
sp. nov.
inhabits lowland evergreen forest on
Penang
Island.
Distribution.
Known only from the
type
locality on
Penang
Island, west
Malaysia
.