Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from Southeast Asia (Araneae, Zodariidae) 3369 Author Dankittipakul, Pakawin Author Jocqué, Rudy Author Singtripop, Tippawan text Zootaxa 2012 2012-07-04 3369 1 1 327 https://biotaxa.org/Zootaxa/article/view/zootaxa.3369.1.1 journal article 10.11646/zootaxa.3369.1.1 1175­5334 5252563 Mallinella platycera sp. nov. ( Figs 158–159 , 162–163 , 169 , 174–185 ) Material examined. Holotype : , THAILAND , Chiang Mai Province : Fang District , Mae Fang NP , Doi Pha Hom Pok , moist evergreen hill forest along a gully ( 2,000–2,200 m ) near the summit ( 2,285 m ), leaf litter sample, 16–17 July 2002 , leg. P. Dankittipakul & P.J. Schwendinger ( MHNG , TH–8A) . Paratypes : 1♂ , 3♀ , same data as holotype ( MHNG , TH–8B). THAILAND , Chiang Mai Province : Chiang Mai City , Doi Suthep-Pui NP : 1♂ , dry deciduous dipterocarp forest, 350 m , 9 May 1986 , leg. P.J. Schwendinger ( MHNG , TH–8C) ; 2♂ , Monthatharn Waterfall, mixed deciduous dipterocarp forest along a gully, 650 m , leaf litter sample, 24 June 2001 , leg. P. Dankittipakul ( MHNG , TH–8D); 1♂ , 600 m , pitfall trap , 15 July 2002 , leg. D. Thapanya ( MHNG , TH–8E) ; 1♀ , evergreen hill forest, 960 m , 23 April – 2 June 1986 ( MHNG ) ; 4♂ , 1♀ , ibidem, 28 April – 30 May 1987 ( TNHM ) ; 1♂ , 2♀ , ibidem, 26 March – 30 July 1986 ( MHNG ) ; 3♂ , ibidem, 2–27 July 1986 ( MHNG ) ; 1♂ , ibidem, 4–28 April 1987 ( TNHM ) . All latter leg. P.J. Schwendinger. 1♀ , Chiang Dao District , Doi Chiang Dao , 510 m , 22 September – 25 October 1990 , leg. P.J. Schwendinger ( MHNG TH–8W) ; 1♀ , outside Tham Klaeb , 3 March 1985 , forest litter, leg. C. Deeleman-Reinhold ( RMHN ) . Chiang Rai Province : 1♂ , Phan District , Doi Luang NP , Poo Kaeng Waterfall , mixed deciduous dipterocarp forest near a waterfall, 800 m , 10 May 2002 , leg. P. Dankittipakul ( MHNG , TH–8F) ; 1♀ , Mae Sai District , Doi Tung , evergreen hill forest, 1,350 m , 30 October 1991 , leg. P.J. Schwendinger ( MHNG , TH–8X) . FIGURES 156–161. 156. Mallinella cryptocera sp. nov. , holotype. 157. M. cryptocera sp. nov. , paratype. 158. M. platycera sp. nov. , holotype. 159. M. platycera sp. nov. , paratype. 160. M. microcera sp. nov. , holotype. 161. M. callicera sp. nov. , holotype. 156–161. Habitus, dorsal. Diagnosis. Males of M. platycera sp. nov. are easily recognized by the second RTA ( Figs 176–178, 179 ) which is relatively broad at base, in combination with the embolus carrying a subterminal process and a triangular apical flange ( Fig. 181 ). Females are distinguished from those of M. cryptocera sp. nov. by a deeper anterior median incision on the epigynal plate and by the narrower apex of their spermathecae in anterior view. Etymology. The specific epithet is a latinized combination of the Greek words (πλατʋς = broad, flat; κƐρας = horn, antler) meaning ‘wide horn’, and refers to the broadest second RTA found among males of this species-group. Noun in apposition. Description. Male ( holotype ). Total length 5.69; prosoma 2.80 long, 2.29 wide; opisthosoma 2.89 long, 2.12 wide. Eye sizes and interdistances: AME 0.13, ALE 0.14, PME 0.15, PLE 0.14, AME–AME 0.14, AME–ALE 0.24, PME–PME 0.18, PME–PLE 0.32; MOQ: 0.50 long, 0.36 anterior width, 0.46 posterior width. Leg measurements: I 7.73 (2.12, 2.38, 1.70, 1.53), II 7.56 (1.95, 2.38, 1.87, 1.36), III 7.48 (2.04, 2.29, 1.95, 1.19), IV 9.11 (2.12, 2.97, 2.80, 1.20). Coloration ( Fig. 158 ). Carapace uniform orange-brown. Sternum reddish brown. Chelicerae brown. Coxae pale yellowish, other leg segments yellowish brown. Dorsum of opisthosoma sepia, mottled with numerous minute pale spots. Dorsal pattern: first and second pairs fused, forming reniform patches; third and fourth pairs represented by two pairs of oblong patches; from fifth pair onwards by a series of transverse median bands. Dorsal scutum orange-brown, edge sharply defined. Palp ( Figs 176–181 ). RTA digitiform, elongated, lateral margins parallel in retrolateral view, apex blunt. Second RTA more or less quadrangular, broad at base, abruptly tapered to rounded apex. Meso-retrolateral side of tibial moderately elevated. TA rostrated, strongly concave prolaterally; with straight apical ridge; apico-prolateral fold sharply pointed, directed mesad; apico-prolateral process absent; baso-retrolateral flange triangular, apex blunt, directed mesad, clearly visible in prolateral view. Conductor terminally sharp and pointed. Embolus flattened, originating at 270°, with broad, semi-transparent flange and lanceolate apex. FIGURES 162–165. 162–163. Mallinella platycera sp. nov. , holotype. 164. M. cryptocera sp. nov. , holotype, 165. M. cryptocera sp. nov. , paratype. 162. Anterior region of carapace. 163. Sternum. 164. PVS, male. 165. Ditto, female. FIGURES 166–169. 166–167. Mallinella cryptocera sp. nov. , holotype. 168. M. cryptocera sp. nov. , paratype. 169. M. platyceras sp. nov. , paratype. 166. Male palp, dorsal. 167. Palpal tibia, retrolateral. 168–169. Epigyne. ERT, elevated retrolateral margin of palpal tibia; RTA, retrolateral tibia apophysis; RTA-2, second retrolateral tibial apophysis. Female ( paratype , MHNG TH–8B). Total length 7.36; prosoma 3.49 long, 2.30 wide; opisthosoma 3.86 long, 2.26 wide. AME 0.13, ALE 0.14, PME 0.15, PLE 0.14, AME–AME 0.14, AME–ALE 0.25, PME–PME 0.14, PME–PLE 0.32; MOQ: 0.52 long, 0.38 anterior width, 0.48 posterior width. Leg measurements: I 7.63 (2.20, 2.30, 1.93, 1.19), II 8.03 (2.17, 2.53, 1.93, 1.39), III 7.67 (2.22, 2.44, 1.80, 1.19), IV 8.22 (2.30, 2.37, 2.20, 1.34). Coloration ( Fig. 159 ). Similar to males but slightly larger in size. Anterior legs yellowish; posterior ones yellowish brown. Genitalia ( Figs 169 , 182–185 ). Epigynal plate rectangular, anterior margin with U-shaped median incision, posterior margin straight. Spermathecae with digitiform apex. Natural history. Mallinella platycera sp. nov. occupies a wide range of habitats, from disturbed, lowland dry dipterocarp forests to pristine evergreen hill forests. Distribution. Widely distributed in northern Thailand .