The millipede genus Tasmaniosoma Verhoeff, 1936 (Diplopoda, Polydesmida, Dalodesmidae) from Tasmania, Australia, with descriptions of 18 new species
Author
Mesibov, Robert
Queen Victoria Museum and Art Gallery, Launceston, Australia
text
ZooKeys
2010
2010-03-26
41
4
31
80
journal article
10.3897/zookeys.41.420
ea2c4da4-3899-41cd-80f4-b7df9302da79
1313–2970
576649
FC5CFE57-05F9-4685-BC02-BB82AB9E4894
Tasmaniosoma
warra
sp. n.
urn:lsid:zoobank.org:act:
C2EDE9C4-E146-4F5B-A848-74068E27C92B
Figs 3A
,
22
; map
Fig. 24
Holotype
.
Male
,
Australia
,
Tasmania
,
Huon River
(Manuka Road),
43°05'36"S
146°43'03"E
(
DN
769286
) ±
100 m
,
100 m
,
29 April 1997
,
R
. Mesibov, plot 1
R4
,
QVM 23
:46037.
Paratypes
.
1 male
, details as for
holotype
,
QVM 23
:46038
;
1 female
, details as for
holotype
but
43°05'42"S
146°42'41"E
(
DN
764284
) ±
100m
,
110 m
, plot 1
R1
,
28 April 1997
,
QVM 23
:46587
;
1 female
, same details but
43°05'49"S
146°42'28"E
(
DN
761282
) ±
100m
,
80 m
,
14 May 1997
, plot 1M6,
QVM 23
:46588
;
1 male
, same details but
43°05'36"S
146°42'45"E
(
DN
765286
) ±
100m
,
130 m
,
15 May 1997
, plot 1
R5
,
QVM 23
:46039
;
1 female
, same details but
43°05'33"S
146°42'41"E
(
DN
764287
) ±
100m
,
140 m
, plot 1
R6
,
QVM 23
:46589
;
1 female
,
Huon River
(
Arve Road
),
Tas
,
43°5'56"S
146°44'45"E
(
DN
792280
) ±
100m
,
170 m
, plot 3M4,
20 May 1997
,
R
.
Mesibov
,
QVM 23
:46590
.
Other
material examined.
12 males
and
10 females
from 16 other sites (see Appendix).
Diagnosis
.
Metatergites without tubercles; ring 6 sternite with discrete setal brushes on anterior margin; gonopod telopodite with stout, rod-like setae in longitudinal tract on posteromedial surface and two large, upright (distally directed) processes arising anterolaterally near apex, the taller, more lateral one Y-shaped, the more anterior one with broad, minutely dentate apical margin.
Description
.
Male/female approximate measurements: length 16/
16 mm
, ring 12 paranota width 1.8/
2.2 mm
. Male as for
T. armatum
in live colour, colour in alcohol and most non-gonopod details, but overall widths tergite 6>5>4>(3,2,head)>collum, ring 12 paranota 1.3 × as wide as prozonite, 6–7 small setae in transverse rows on midbody metatergites (
Fig. 3A
), ozopore opening very close to paranotal margin. Ring 6 sternite as for
T. armatum
, but with paired paramedian brushes of fine setae on anterior margin separated by shallow notch.
Gonopod telopodite (
Fig. 22
) straight, slightly constricted just above base, with narrow buttress near base on anterolateral surface. Divided between 2/3 and 3/4 telopodite height into five processes (noted here lateral to medial): (a) large, anteroposteriorly flattened process, widely expanded distally, distal margin lower medially and minutely dentate, lateral end of margin broadly acuminate; (b) large, Y-shaped process, flattened anteroposteriorly, divided at ca 2/3 process height, tips roundly acuminate; (c) mediolaterally flattened laminate process, hook-shaped posteriorly; (d) solenomere with slender, distally directed, posteriorly curving distal branch and short, broad, posteriorly directed basal branch; (e) small, slender medial process arising basal to solenomere origin, directed posterodistally, with expanded, truncate tip. Posterodistally directed tract of 20–30 stout, rod-like setae on posteromedial surface from ca 1/3 to 2/3 telopodite height; small group of fine setae on posterolateral surface just above base; longitudinal row of 5–6 fine setae laterally at mid-height on telopodite. Prostatic groove running distally and slightly posteriorly on posteromedial surface to base of solenomere.
Figure 22.
Tasmaniosoma warra
sp. n.
, paratype male, QVM 23:46038
A
and voucher male QVM 23:46040
B–D
.
A
Right posterolateral view of gonopods in situ
B
medial,
C
anterior and
D
lateral views of right gonopod telopodite. Setation not shown in
B–D
; dashed line marks course of prostatic groove. Scale bars = 0.2 mm.
Female more robust than male; posterior margin of epigynum slightly elevated at both ends and produced medially as small, rounded triangle.
Distribution
.
Common in wet eucalypt forest (with and without a cool temperate rainforest understorey) over ca
2500 km
2
on the southeast Tasmanian mainland and on South Bruny Island, from near sea level to at least
370 m
(
Fig. 24
). On the Tasmanian mainland, not yet found more than a few kilometres north of Grove, south of Dover or west of Mt Weld. Co-occurs with
T. bruniense
sp. n.
on Bruny Island; possibly parapatric with
T. australe
sp. n.
, as the two species occur within ca
7 km
of each other and have not yet been found together.
Etymology
.
Noun in apposition, for the
type
locality in the Warra Long Term Ecological Research (LTER) site in the Southern Forests.