Description of a Cretaceous amber fossil putatively of the tribe Coprophilini (Coleoptera, Staphylinidae, Oxytelinae) Author Makranczy, Gyoergy Author Yamamoto, Shuhei Author Engel, Michael S. text ZooKeys 2018 782 81 94 http://dx.doi.org/10.3897/zookeys.782.27733 journal article http://dx.doi.org/10.3897/zookeys.782.27733 1313-2970-782-81 789CE1ADE7784C898DBFAD7A01C7ECDE 789CE1ADE7784C898DBFAD7A01C7ECDE Gollandia planata sp. n. Figures 1-2, 3-4, 5-6, 7-11 Holotype. Sex unknown, probably male, in a flattened drop shaped, light yellow amber piece (20.0 x 9.9 x 4.5 mm, 0.98g): "FMNHINS 3729858 ex S. Yamamoto collection (SYAC0482)" deposited in Field Museum of Natural History (Chicago, USA). Locality and horizon. Noije Bum hill near Tanai Village, SW part of Hukawng Valley (SW of Maingkhwan), Kachin State, northern Myanmar; lowermost Cenomanian, Upper Cretaceous. Diagnosis. As for the genus (vide supra). Description. Measurements: HW = 0.45; TW = 0.41; PW = 0.64; SW = 0.59; MW = 0.68; AW = 0.70; HL = 0.29; EL = 0.10; TL = 0.04; PL = 0.50; SL = 0.66; SC = 0.55; FB = 1.47; BL = 3.29 mm (all measured from dorsal view). Habitus: General habitus as in figures 1-6. Colour reddish dark brown. Body moderately lustrous, covered with fine microsculpture and forebody finely, not very densely setose. Abdomen with longer and stronger lateral setae posteriorly. Head. Head rather short. Antennae rather elongate, scape almost twice as wide as pedicel and not much longer, second antennomere (pedicel) more than 3.5 x as long as wide, third antennomere (first flagellomere) slender at base and almost as long as previous. Further antennomeres spindle-shaped and each with rudimentary basal dish, gently constricted above them. Antennomeres 4-7 at least 2.5 x as long as wide, from antennomere 8 becoming wider, gently clubbed, last three antennomeres only about 1.5 x as long as wide. Compound eyes more than 2 x as long as weakly formed temples. Neck not constricting strongly. Thorax. Pronotum rather large, widest point slightly before middle with both anterior and posterior corners rather narrowly rounded, lateral margin slightly concave before quite acute posterior angles. Surface finely microsculptured, thereby punctation partly obscured. Disc transversally impressed before base (in a curved fashion), also with a semi-triangular mid-longitudinal impression anteriad; rather large but shallow paralateral depressions on sides. Elytra. Elytra together just slightly broader than pronotum, trapezoidal, shoulders well developed, narrowly rounded, even slightly projecting forward in relation to anterior edge at mesoscutellar area. Dorsal surface finely punctate and setose, no major lateral setae, epipleural ridge with moderately long setae at regular intervals. Abdomen. Sides of abdomen gently curved, almost parallel. Surface of tergites with moderately fine, longitudinally elongate punctures, apical edges of tergites (up to tergite VI) with row of equal-sized setae at regular intervals. Specimen is without any feature suggesting strong sexual dimorphism. No genital traits observable. Figures 1-2. Gollandia planata gen. et sp. n. 1 habitus, dorsal view 2 sketch of main body parts, ventral view. Figures 3-4. Gollandia planata gen. et sp. n. photographed with macro lens and three flash units. 3 dorsal view 4 ventral view. Figures 5-6. Gollandia planata gen. et sp. n. photographed through microscope, strong backlighting. 5 dorsal view 6 ventral view. Etymology. The specific epithet is a Greek adjective derived from platys (= wide) and refers to the pronotum of the species being unusually explanate. Preservation. The specimen is exceedingly well preserved, with the hind wings unfolded over part of the abdomen, minor air bubbles under segmental edges, a thin air layer over some sculptured dorsal parts, and the ventral side exceptionally clearly visible. As explained before, the specimen is sitting within a layer of resin covered by another layer, and this creates an effect similar to the specimen being glued to a glass, evident in the photos of the ventral side. The legs are somewhat distorted (but each pair is almost perfect on one side). Primitive oxytelines often have distinctive coxites and styli in females (if not exposed, then setation gives them away), and in their absence the specimen is presumed to be a male. Figures 7-11. Gollandia planata gen. et sp. n. 7 head, pro- and mesothorax, ventral view 8 fore-body, dorsal view 9 meso- and metacoxae, ventral view 10 mid leg, dorsal view 11 protarsus, ventral view. t 1-4 = tarsomeres, em = empodial seta, ts = mesotibial spur, sc = mesoscutellar impression, ec = elytral posterior corner, gs = gular sulcus, pf = procoxal fissure, ms = mesosternal, and mt = metasternal process. Scale bars: 0.15 mm.