Six new species of Australocosmica Köhler, 2011 from the Kimberley islands, Western Australia (Mollusca: Pulmonata: Camaenidae)
Author
Criscione, Francesco
Author
Köhler, Frank
text
Zootaxa
2013
3608
2
101
115
journal article
10.11646/zootaxa.3608.2.1
dc20b97c-cdf4-467d-8ce9-98e912fe060c
1175-5326
223235
0E56C847-5694-4315-9A51-B4F207CA4F91
Australocosmica nana
n. sp.
Figs 3
D, 5A–B, 6D
Holotype
.
Australia
, WA, Kimberley, Bonaparte Archipelago, Albert Island: Sta. KC-043, Bonaparte Archipelago, -14.519, 124.929 (coll. V.
Kessner, A.
Longbottom, 14 July
1988
)
(WAM
S84060
).
Paratypes
.
Australia
, WA, Kimberley, Bonaparte Archipelago, Albert Island: same data as for
holotype
, 14 wet (WAM
S83039
); 36 dry (WAM
S83038
)
,
35
dry (FMNH
219165
).
Etymology
. In reference to its small shell, derived from “
nanus
” (Latin=dwarf), adjective of feminine gender.
FIGURE 6.
Penial anatomy of
Australocosmica
species.
A.
A. bernoulliensis
n. sp.
(holotype WAM
S84059
).
B.
A. buffonensis
n. sp.
(holotype WAM
S84059
).
C.
A. crassicostata
n. sp.
(holotype WAM
S84059
).
D.
A. nana
n. sp.
(holotype WAM
S84060
). Scale bar = 2 mm.
FIGURE 7.
Penial anatomy of
Australocosmica
species.
A.
A. pallida
n. sp.
(holotype WAM
S84056
).
B.
A. rotunda
n. sp.
(holotype WAM
S84061
). Scale bar = 2 mm.
FIGURE 8.
Map showing distributions of
Australocosmica
in the Kimberley (WA). Insets:
A.
Maret Island group with
A. crassicostata
n. sp.
(․) and
A. nana
n. sp.
(●).
B.
Coronation island group with
A. bernoulliensis
n. sp.
(♦) and
A. buffonensis
n. sp.
(˔).
C.
central Kimberley coast, Prince Regent Reserve with
A. pallida
n. sp.
(♣) and
A. rotunda
n. sp.
(♠).
Description.
Shell (
Figs 3
D; 5A, B). Small, with short spire and sub-globose shape. Whorls with slightly angulate periphery, moderately shouldered underneath suture. Axial ribs moderately pronounced. Colour horn to light brown. Genitalia (
Fig. 6
D). Penis about half as long as anterior part of oviduct (= vagina and free oviduct). Inner penial wall supporting complex sculpture, with series of folds surrounding the opening to vas deferens forming a collar, posteriorly with three corrugated, wide, longitudinal pilasters, anteriorly with numerous, narrow longitudinal pilasters extending into atrium. Vaginal length equivalent to ¾ of penis and free oviduct. Spermoviduct 3.5 times longer than free oviduct and almost twice as long as albumen gland.
Remarks:
Shell colour as variable as
A. crassicostata
. Shell smaller, flatter and with fewer whorls than other congeners. Sculpture similar to that of
A. rotunda
and with fewer but more pronounced axial ribs than other congeners except
A. crassicostata
. Inner penial wall sculpture similar to that of
A. buffonenis
and
A. pallida
but with more developed transverse folds on lateral pads. This is Solem’s (1991) manuscript species ‘NSP 80’.