Order Scandentia Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 104 109 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Tupaia moellendorffi Matschie 1898 Tupaia moellendorffi Matschie 1898 , Sitzb. Ges. Naturf. Fr. Berlin: 39 . Type Locality: Philippines , Culion Isl. Vernacular Names: Calamian Treeshrew . Subspecies: : Subspecies Tupaia moellendorffi subsp. moellendorffi Matschie 1898 Subspecies Tupaia moellendorffi subsp. busuangae Sanborn 1952 Subspecies Tupaia moellendorffi subsp. cuyonis Miller 1910 Distribution: Calamian Isls (Busuanga, Culion) and Cuyo in the Philippines . Conservation: CITES – Appendix II; IUCN – Vulnerable (as included in T. palawanensis ). Discussion: The three named forms included here appear more closely-related to each another than any is to T. palawanensis (where they are usually arranged; e.g. Heaney et al., 1998 ) and I provisionally separate them here as a distinctive complex; see also comments by Lyon (1913) . This arrangement is somewhat more consistent with species-boundaries traditionally recognized in squirrels from the Palawan region (see accounts of Sundasciurus ).