<document ID-CLB-Dataset="25968" ID-DOI="http://dx.doi.org/10.3897/zookeys.872.35361" ID-GBIF-Dataset="ed1a1322-b118-4434-b9ff-503a0fb64792" ID-PMC="PMC6711934" ID-Pensoft-Pub="1313-2970-872-13" ID-Pensoft-UUID="A9BE568D9BF65A179464AB8C7CB8BEB2" ID-PubMed="31496884" ID-ZooBank="DCDEFCB1E33D476F90D7178BC08C311A" ModsDocID="1313-2970-872-13" checkinTime="1566364211746" checkinUser="pensoft" docAuthor="Astafurova, Yulia V., Proshchalykin, Maxim Yu. &amp; Schwarz, Maximilian" docDate="2019" docId="7CC2FBA91AFE532A829B9A3365B2C600" docLanguage="en" docName="ZooKeys 872: 13-40" docOrigin="ZooKeys 872" docSource="http://dx.doi.org/10.3897/zookeys.872.35361" docTitle="Sphecodes intermedius Bluethgen 1923" docType="treatment" docVersion="6" id="A9BE568D9BF65A179464AB8C7CB8BEB2" lastPageNumber="23" masterDocId="A9BE568D9BF65A179464AB8C7CB8BEB2" masterDocTitle="The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010" masterLastPageNumber="40" masterPageNumber="13" pageNumber="22" updateTime="1732684065680" updateUser="ExternalLinkService">
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<mods:title id="4E36844052C19AC1F6F6E22D0721F14C">The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010</mods:title>
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<mods:namePart id="A85A764BE02A9DE01D7E54CAC15735FE">Astafurova, Yulia V.</mods:namePart>
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<mods:namePart id="C5ED7D69A9C2207648F8975F04436973">Proshchalykin, Maxim Yu.</mods:namePart>
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<mods:namePart id="594782A9B7BFC8264C657A5F4227BA4C">Schwarz, Maximilian</mods:namePart>
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<mods:date id="9B3497237B0DE1F849D797F2F3D2E51B">2019</mods:date>
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<mods:classification id="E1D6F8BA71FB7277F73D20EE91A5A0CE">journal article</mods:classification>
<mods:identifier id="31FC3AF317CAC7104474BC2B8EC36127" type="DOI">http://dx.doi.org/10.3897/zookeys.872.35361</mods:identifier>
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<treatment id="7CC2FBA91AFE532A829B9A3365B2C600" ID-GBIF-Taxon="159217133" LSID="urn:lsid:plazi:treatment:7CC2FBA91AFE532A829B9A3365B2C600" httpUri="http://treatment.plazi.org/id/7CC2FBA91AFE532A829B9A3365B2C600" lastPageId="10" lastPageNumber="23" pageId="9" pageNumber="22" scope_family="Halictidae" scope_order="Hymenoptera">
<subSubSection id="BD4AA8ED9106C5D9BB147280135DD805" pageId="9" pageNumber="22" type="nomenclature">
<paragraph id="B69BE924FFEFB8637DE29389C187E3E4" pageId="9" pageNumber="22">
<taxonomicName id="E5D0F9353F1EDEE5AF8CBEAA1796A331" ID-CoL="6Z9N8" LSID="7cc2fba9-1afe-532a-829b-9a3365b2c600" authority="Bluethgen, 1923" authorityName="Bluethgen" authorityYear="1923" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes intermedius" order="Hymenoptera" pageId="9" pageNumber="22" phylum="Arthropoda" rank="species" species="intermedius">
<pageBreakToken id="388AA7EFF08BDF5193C06FC31DBFE0AC" pageId="9" pageNumber="22" start="start">Sphecodes</pageBreakToken>
intermedius 
<normalizedToken id="B740F045B9120C0FBBDD5A934329888C" originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1923
</taxonomicName>
<figureCitation id="CF03E2F82EC6398C6401D5CBB7F7F5CE" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="9" pageNumber="22">Figure 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A7C189316992430FF6B579EDED4B7C41" pageId="10" pageNumber="23" type="reference_group">
<paragraph id="513E180251B8E33AE612296C88E5AE30" pageId="10" pageNumber="23">
<taxonomicName id="970C95028E0A72C898BEA26B4290D377" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes intermedius" order="Hymenoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="intermedius">
<pageBreakToken id="F4FF3732BA57216C17E3563A5C9A7529" pageId="10" pageNumber="23" start="start">Sphecodes</pageBreakToken>
intermedius
</taxonomicName>
<normalizedToken id="AB8EE216845AB89930F9836A99E4DF82" originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1923: 500 (lectotype (
<emphasis id="63934B499F3B5B7F274F43687DAC07E5" bold="true" pageId="10" pageNumber="23">designated here</emphasis>
): ♂, Type &lt;red label&gt; // Caucas Portz // Sph. 
<taxonomicName id="05B502BF68260004E32A32EC9BBC7B6C" class="Insecta" family="Halictidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="intermedius" order="Hymenoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="intermedius">intermedius</taxonomicName>
♂, Type., P. 
<normalizedToken id="43758C09DA6C1A7FF74D920EA6867EE4" originalValue="Blüthgen">Bluethgen</normalizedToken>
det. // &lt;blue circle&gt; // Lectotypus, 
<taxonomicName id="EFDC4D35F34780D7362BBAAC6F890B7A" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes intermedius" order="Hymenoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="intermedius">Sphecodes intermedius</taxonomicName>
<normalizedToken id="14FAF4B881D4B99D2152BEEA3CAE4532" originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1923, ♂, des. Astafurova &amp; Proshchalykin, 2018; paralectotype: ♀, Type &lt;red label&gt; Sph. 
<taxonomicName id="8F402AAB160AB6644C8926D84F385B52" class="Insecta" family="Halictidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="intermedius" order="Hymenoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="intermedius">intermedius</taxonomicName>
♀, Type, P. 
<normalizedToken id="61812FD73F99355EF357A98CBEDB7B97" originalValue="Blüthgen">Bluethgen</normalizedToken>
det. // Paralectotypus, 
<taxonomicName id="5D064A07C4AAA3166EE2ADC81F212A54" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes intermedius" order="Hymenoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="intermedius">Sphecodes intermedius</taxonomicName>
<normalizedToken id="7BDA8210356619F704CB378BBFC5A0CD" originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1923, ♀, des. Astafurova &amp; Proshchalykin, 2018; ISZP, examined, 
<figureCitation id="C2292FC7EDC679171CEE189DAED4A935" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="10" pageNumber="23">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="54E55A09AF1174FC34B5CB2B78BE95F9" pageId="10" pageNumber="23">
<taxonomicName id="64082DC088FE9C7FC69A14FB68BB7F25" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes lactipennis" order="Hymenoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="lactipennis">Sphecodes lactipennis</taxonomicName>
Meyer, 1925 (Synonym).
</paragraph>
</subSubSection>
<subSubSection id="AE3FBBF342E83AB6404C371EC1E1DF06" pageId="10" pageNumber="23" type="diagnosis">
<paragraph id="ACDAD70B1FD850960B34F770A5491A0C" pageId="10" pageNumber="23">Diagnosis.</paragraph>
<paragraph id="3181A2991229A331707E7DBB760AE359" pageId="10" pageNumber="23">
See 
<bibRefCitation id="AC1DFA3275E85B46A5C9D8849C43CDED" DOI="https://doi.org/10.25221/fee.369.1" author="Astafurova, YuV" journalOrPublisher="Far Eastern Entomologist" pageId="22" pageNumber="35" pagination="1 - 47" refId="B13" refString="Astafurova, YuV, Proshchalykin, MYu, Engel, MS, 2018a. The cuckoo bee genus Sphecodes Latreille, 1804 in Kazakhstan (Hymenoptera: Halictidae). . Far Eastern Entomologist 369: 1 - 47" title="The cuckoo bee genus Sphecodes Latreille, 1804 in Kazakhstan (Hymenoptera: Halictidae)." url="https://doi.org/10.25221/fee.369.1" volume="369" year="2018 a">Astafurova et al. 2018a</bibRefCitation>
: 20.
</paragraph>
</subSubSection>
<subSubSection id="3E93AFE251ABA5E170EEA261DC18BCC4" pageId="10" pageNumber="23" type="materials_examined">
<paragraph id="A1AECC10EAA276CB307A46B2AD9AEC77" pageId="10" pageNumber="23">Material examined.</paragraph>
<paragraph id="476BD6F7B9217CA25A13DF0C25FA56AF" pageId="10" pageNumber="23">
UNITED ARAB EMIRATES: 1 ♂, Hatta (Hotel), 28. IV.1989, (NHMUK 013380370); 1 ♂, idem, 23.VIII.1991 (NHMUK 013380409); 1 ♂, idem, 14.IV.1990, I. L. Hamer [D. Baker det., 1992 as 
<taxonomicName id="FFD7CA40DB56A8231E86DD8103428167" lsidName="S. punctatissimus" pageId="10" pageNumber="23" rank="species" species="punctatissimus">
<emphasis id="7C4EF72FCA41CCB84AA5548B75A4EE2A" italics="true" pageId="10" pageNumber="23">S. punctatissimus</emphasis>
</taxonomicName>
Meyer] (NHMUK 013380361); ISRAEL: 1 ♀, Jerusalim, 16.VII.1930, S. Bodenheimer [det. 
<normalizedToken id="1CAEEBF2096C6A6C05BF3DDC7D679F4A" originalValue="Blüthgen">Bluethgen</normalizedToken>
] (MNHB); 1 ♀, Tiberias, 200 m, 22.III.1975, K.M. Guichard (NHMUK 013380410); 1 ♀, Jericho (Hisham Palace), 200 m, 8.III.1975, K.M. Guichard (NHMUK 1975-248, 013380408); JORDAN: 1 ♀, N. Shuna env., 20-22.IV.1996; 1 ♀, idem, 29-30.IV.1996, M. Halada (
<normalizedToken id="5E27A8938A1DEC063D15C8857C00DC3A" originalValue="OÖLM">OOELM</normalizedToken>
)
</paragraph>
</subSubSection>
<subSubSection id="432925BF6E0A5B298B61FA08050EE3CB" pageId="10" pageNumber="23" type="published records">
<paragraph id="D2BEF5E54F7453D87E04A293CDC41B53" pageId="10" pageNumber="23">Published records.</paragraph>
<paragraph id="7421644B20ADE03BCDF52067CFC4BFA2" pageId="10" pageNumber="23">
<bibRefCitation id="0ED378E7A95CEB0F76CB44B647BA6C93" author="Ascher, JS" journalOrPublisher="Zoologica Scripta" pageId="21" pageNumber="34" publicationUrl="http://www.discoverlife.org/mp/20q?guide=Apoidea_species" refId="B3" refString="Ascher, JS, Pickering, J, 2019. . http://www.discoverlife.org/mp/20q?guide=Apoidea_species" url="http://www.discoverlife.org/mp/20q?guide=Apoidea_species" year="2019">Ascher and Pickering 2019</bibRefCitation>
(Israel)
</paragraph>
</subSubSection>
<subSubSection id="800273FD4FD6B14A364A0ECD9634D3B6" pageId="10" pageNumber="23" type="distribution">
<paragraph id="89F68B2DBF3059B6DEBFC3EE2DB5836B" pageId="10" pageNumber="23">Distribution.</paragraph>
<paragraph id="23E8646D2DCB64B59DCD7C188C7BC92F" pageId="10" pageNumber="23">*United Arab Emirates, Israel, *Jordan; North Africa, South Europe (east to Ukraine), Russia (south of the European part, Urals), Caucasus, Turkey, Kazakhstan, Central Asia, Pakistan, China (Gansu).</paragraph>
</subSubSection>
<subSubSection id="F8C0083A0BBA7E222C35D8ADEC49ADB6" pageId="10" pageNumber="23" type="remarks">
<paragraph id="718215A5B892CE710D867F3E6F08908C" pageId="10" pageNumber="23">Remarks.</paragraph>
<paragraph id="BE6DFC15806BA37B40D63FE7F3191855" pageId="10" pageNumber="23">
<taxonomicName id="578497CB81A61122A42BBF3364F7E800" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes intermedius" order="Hymenoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="F002DFD588362C9E05DD8405669703F0" italics="true" pageId="10" pageNumber="23">Sphecodes intermedius</emphasis>
</taxonomicName>
<normalizedToken id="F4AC0A2C44A4683CFA31D24DB4655CF5" originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1923 was described from specimens of both sexes collected in 
<normalizedToken id="0D5FF93B7A0B3A1CFCEA5B9AE321481D" originalValue="“Caucas”">&quot;Caucas&quot;</normalizedToken>
[Caucasus] (
<figureCitation id="95ABA6A861B580DAFFA391346C1A9542" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="10" pageNumber="23">Fig. 11</figureCitation>
). There are two specimens (female and male) in ISZP from this locality, which correspond to the original description of P. 
<normalizedToken id="0B8557867A15BBC2A2E8652FF4B109B2" originalValue="Blüthgen">Bluethgen</normalizedToken>
. One of these specimens (male) is designated here as a lectotype of 
<taxonomicName id="CCDDB25E5E2279E4549C3F855D8C1367" lsidName="S. intermedius" pageId="10" pageNumber="23" rank="species" species="intermedius">
<emphasis id="5E1987A633FE0BC79FA3A1822FEDDB5E" italics="true" pageId="10" pageNumber="23">S. intermedius</emphasis>
</taxonomicName>
to avoid any confusion about the status of the type specimens and to properly diagnose this species.
</paragraph>
</subSubSection>
</treatment>
</document>