An update on the distribution of the poorly known flower beetle genus Cymophorus Kirby, 1827 (Coleoptera: Scarabaeidae: Cetoniinae) in Asia
Author
Xu, Hao
Author
Qiu, Jian-Yue
text
Zootaxa
2020
2020-11-11
4877
3
575
586
journal article
8013
10.11646/zootaxa.4877.3.9
a5e1f244-a812-43da-a18b-05e417571c56
1175-5326
4424497
DF4B0414-D99D-4D6B-8C36-E03FDECC099D
Cymophorus
Kirby, 1827
Cymophorus
Kirby 1827: 153
;
Schaum 1845: 53
(in synonymy);
Schaum 1848: 60
(in synonymy);
Lacordaire 1856: 544
(in synonymy); Harold in
Gemminger & Harold 1869: 1334
(catalogue);
Westwood 1873: 16
;
Bergé 1884: 157
;
Schoch 1894: 218
(in key);
Schoch 1895: 132
;
Schoch 1896: 78
(catalogue);
Péringuey 1907: 499
(South African species);
Arrow 1910: 203
(Indian species);
Schenkling 1921: 358
(catalogue);
Schein 1954: 357
(revision);
Paulian 1961: 4
;
Krikken 1977: 310
(in key);
Krikken 1984: 44
;
Marais & Holm 1992: 20
(southern African species);
Ma 1995: 150
(
China
);
Sakai & Nagai 1998: 153
;
Krajčík 1999: 35
(catalogue);
Krajčík 2011: 68
;
Krajčik 2012: 85
(list);
Bezděk 2016: 391
(list).
Type
species
Cymophorus undatus
Kirby, 1827
, by monotypy.
Cymatophorus
Burmeister in
Burmeister & Schaum, 1840: 361
(unjustified emendation);
Schenkling 1921: 358
(in synonymy);
Marais & Holm 1992: 20
;
Krajčík 1999: 35
(in synonymy).
Ptychophorus
Schaum in
Burmeister & Schaum 1841: 271
(unnecessary replacement name);
Burmeister 1842: 646
;
Schaum 1845: 53
(catalogue);
Schaum 1848: 60
(catalogue);
Lacordaire 1856: 544
; Harold in
Gemminger & Harold 1869: 1334
(in synonymy);
Westwood 1873: 16
(in synonymy);
Péringuey 1907: 499
(in synonymy);
Arrow 1910: 203
(in synonymy);
Schenkling 1921: 358
(in synonymy);
Paulian 1961: 4
(in synonymy);
Marais & Holm 1992: 70
(in synonymy);
Ma 1995: 150
(in synonymy);
Krajčík 1999: 35
(in synonymy);
Bezděk 2016: 391
(in synonymy).
Nomenclature.
The name
Cymophorus
has not been constantly used in early times that produced two junior synonyms. Schaum claimed that the generic name was preoccupied by
Cymatophora
Treitschke, 1825
(
Lepidoptera
:
Thyatiridae
), and introduced the name
Ptychophorus
as a replacement for this genus (
Burmeister & Schaum 1841
). This replacement name was adopted in a few subsequent works (
Burmeister 1842
,
1845
;
Schaum 1844
,
1845
,
1848
). Later
Lacordaire (1856)
and
Westwood (1873)
pointed out that the spelling of Kirby’s
Cymophorus
is sufficiently different from Treitschke’s
Cymatophora
, and the original generic name should be resurrected. Actually, the replacement name
Ptychophorus
was not proposed for Kirby’s
Cymophorus
, and the spelling of Kirby’s name was already changed to
Cymatophorus
by Burmeister (
Burmeister & Schaum 1840: 361
) without explanation. This modified name appears to be overlooked; nevertheless, Kirby’s
Cymophorus
remains the valid name of this genus, because Article 56.2 of the International Code of Zoological Nomenclature (
ICZN 1999
) indicates that two genus-group names are not homonyms even if the difference between them is only one letter. Hence,
Ptychopho-rus
Schaum, 1841 being an unnecessary substitute name. Besides, according to Article 33.2.3 (
ICZN 1999
),
Cy-matophorus
Burmeister, 1840
is an unjustified emendation, and thus, as
Schein (1954)
stated, it should be treated as a junior objective synonym of
Cymophorus
. This junior synonym was listed in
Krajčík (1999)
but neglected in the new Palaearctic catalogue (
Bezděk 2016
).
Remarks.
When divided the genus
Cymophorus
into seven subgenera,
Schein (1954)
assigned four African species to the subgenus
Pseudocymophorus
Schein, 1954
with the Asian species
C. pulchellus
. However,
Marais & Holm (1992)
listed two of them,
C. sexfoveatus
Fairmaire, 1884
and
C. limbatus
Gestro, 1895
, in the nominate subgenus without explanation. Later
Krajčik (1999
,
2012
) not only followed this arrangement but also incorrectly placed
C. pulchellus
in the nominate subgenus. To confirm the true placement of these species requires a detailed study of other congeners. It is beyond the scope of this study, and thus we adopt the arrangement in
Schein’s (1954)
revision.