A revised classification of the family Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights Author Last, Peter R. Author Naylor, Gavin J. P. Author Manjaji-Matsumoto, B. Mabel text Zootaxa 2016 4139 3 345 368 journal article 10.11646/zootaxa.4139.3.2 51358935-98b0-4d89-a47f-e4341171e5b9 1175-5326 262765 DCF4220B-4A73-407C-837C-54DEAE29F435 Genus Neotrygon Castelnau, 1873 Neotrygon Castelnau, 1873:122 . Type species Raya trigonoides Castelnau, 1873 ; by monotypy. Definition . Small dasyatids (adults to 27–47 cm DW) characterised by the following: moderately depressed to robust, rhombic disc with pectoral-fin apex typically narrowly angular; snout obtuse and short (1.2–2.3 times combined orbit and spiracle length); eye large and protruding (often greatly); nasal curtain narrow, typically subrectangular; mouth narrow with 2 large median oral papillae; tail flexible and short (length 1.1–1.5 times DW), its base medium-broad and depressed to flattened oval; pelvic fins long and pointed, protruding slightly to wellbeyond disc; dorsal fold well developed; ventral fold low with a long base; caudal sting positioned moderately well back on tail (distance from pectoral-fin insertion to caudal-sting base 2.1–2.8 times interspiracular width); skin smooth and denticle band absent; row of small median thorns in most adults, no other thorns on disc; tail smooth; typically with strong dorsal colour pattern, usually with dark mask-like marking around eyes and blue or black spots on disc; posterior tail with black and white bands; ventral surface white, disc margin typically darker; marine, Indo– West Pacific. Species . N. annotata ( Last, 1987 ) , N. australiae Last, White & Séret, 2016 , N. caeruleopunctata Last, White & Séret, 2016 , N. kuhlii (Müller & Henle, 1841) , N. leylandi ( Last, 1987 ) , N. ningalooensis Last, White & Puckridge, 2010 , N. orientale Last, White & Séret, 2016 , N. picta Last & White, 2008 , N. trigonoides (Castelnau, 1873) , and N. varidens (Garman, 1885) . Remarks . The taxonomy of a subgroup of Neotrygon , the kuhlii-complex, is not fully resolved ( Puckridge et al. , 2013 ). Three new members of the kuhlii-complex were described recently and a Western Pacific species ( N. varidens ) was resurrected (Last et al. , 2016). Members of this subgroup are very similar in morphology and an Indo–Pacific revision is needed.