Revisions and key to the Vernonieae (Compositae) of Thailand Author Bunwong, Sukhonthip Maejo University Phrae Campus, Mae Sai, Rong Kwang, Phrae 54140, Thailand Author Chantaranothai, Pranom Applied Taxonomic Research Center, Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand Author Keeley, Sterling C. Department of Botany, University of Hawaii, Honolulu, HI 96816 USA text PhytoKeys 2014 2014-05-13 37 25 101 http://dx.doi.org/10.3897/phytokeys.37.6499 journal article http://dx.doi.org/10.3897/phytokeys.37.6499 1314-2003-37-25 FFE8FFACFF84FFA95573FFFECD03F742 576215 Pseudelephantopus spicatus (Juss. ex Aubl.) C.F. Baker, Trans. Acad. Sci. St. Louis 12: 45, 55 & 56. 1902. Pseudelephantopus spicatus (Juss. ex Aubl.) Rohr, Skrifl. Nat. Selsk. Kiobenh. 2: 216. 1792. Elephantopus spicatus Juss. ex Aublet, Hist. Pl. Guiane 2: 808. 1775. Type. not ascertained. Figs 8E-F . Description. Perennial herbs, 10-40 cm tall. Stems erect, inconspicuously ribbed, puberulous. Leaves simple, rosulate or alternate at base, 5-15 by 1.5-5 cm, obovate or oblanceolate, margin slightly serrate to entire, apex obtuse or rounded, base cuneate or attenuate, subcoriaceous; upper surface puberulous without glands, lower surface puberulous with filiform hairs and capitate glands; lateral veins 9-15-paired; petioles up to 2 cm long. Capitulescences terminal and axillary, capitula 1-4 aggregated in clusters supported by foliaceous bracts, clusters arranged in a spike. Capitula tubular, 14-17 mm long. Receptacle flat, 1-1.5 mm in diam., glabrous. Florets bisexual and fertile. Involucres oblong, in 2 series, 10-11 mm long. Phyllaries 8, decussate, light green, margin entire or piliferous, outer surface pilose without glands; the outer lanceolate, apicies acute; the inner ones obovate-lanceolate or oblong, apices acute. Florets 4; corollas salverform, white, zygomorphic, glabrous; corolla tubes 5-9 mm long; corolla lobes 2.5-2.8 mm long. Anthers 1.5-2 mm long, apical appendage acute, base acute. Styles white, 5-9 mm long, branches ca. 2 mm long, inner surface covered with stigmatic papillae. Achenes clavate, 4-5 mm long, pubescent with densely twin hairs, without glands, 10-ribbed. Pappus in 1 series, often of 2 sizes and bent at the tip, bristles 6-9, 2-6 mm long. Distribution. Thailand: Chiang Rai, Nakhon Phanom, Ubon Ratchathani. Tropics. Specimens examined. Thailand, Ubon Ratchathani, Chong Mek border crossing, 15°8.02'N , 105°28.01'E , 27 Oct 2007, S. Bunwong 342 (KKU); Chiang Rai, Muang District, 20 Nov 2007, S. Bunwong 352 (KKU, US); Nakhon Phanom, Ban Phang, 24 Feb 2003, Th. Wongprasert et al. 032-17 (BKF). Diagnostic characters. Pseudelephantopus spicatus is distinguished from Elephantopus by having spicate capitulescences and contorted pappus. Ecology. Open areas in dipterocarp forest or river bank, alt. 100-400 m; flowering October to December. Vernacular name. Doo La Doo (โด่ลาโด่), Ton Tai Din (ต้นใต้ดิน).