<document id="2404B8CFCF87830E55197F4929AFC05C" ID-DOI="10.5852/ejt.2020.663" ID-GBIF-Dataset="5a7c4049-6179-47df-8537-a81f307a7dc2" ID-ISSN="2118-9773" ID-Zenodo-Dep="3890442" ID-ZooBank="78E7049C-F592-4D01-9D15-C7715119B584" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1591910064537" checkinUser="valdenar" docAuthor="Tenorio, Manuel J., Abalde, Samuel, Pardos-Blas, José R. &amp; Zardoya, Rafael" docDate="2020" docId="AE38686E301EFFCFFE31F886FC540A20" docLanguage="en" docName="ejt-663_tenoria_abalde_pardos-blas.pdf.imf" docOrigin="European Journal of Taxonomy 663 (663)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Varioconus tacomae Tenorio &amp; Abalde &amp; Pardos-Blas &amp; Cmim 2020, comb. nov." docType="treatment" docVersion="13" lastPageNumber="74" masterDocId="520110163059FF86FFF7FF98FFFF091D" masterDocTitle="Taxonomic revision of West African cone snails (Gastropoda: Conidae) based upon mitogenomic studies: implications for conservation" masterLastPageNumber="89" masterPageNumber="1" pageNumber="72" updateTime="1733342609484" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="897BFFD3D851CD9F1DEBE0AF5F157ADE">Taxonomic revision of West African cone snails (Gastropoda: Conidae) based upon mitogenomic studies: implications for conservation</mods:title>
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<mods:namePart id="3829A3C4430DFA65F3D53174F3D235A4">Tenorio, Manuel J.</mods:namePart>
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<mods:affiliation id="EDD9661027DB0E75FBBB55C967FD6E2B">Departamento CMIM y Química Inorgánica - Instituto de Biomoléculas (INBIO), Facultad de Ciencias, Torre Norte, 1 ª Planta, Universidad de Cadiz, 11510 Puerto Real, Cadiz, Spain.</mods:affiliation>
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<mods:namePart id="09162C62B06CA04330DFDA77D68384FB">Abalde, Samuel</mods:namePart>
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<mods:affiliation id="D1EE80985BAC1C9CCAED553105757912">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain.</mods:affiliation>
<mods:nameIdentifier id="20A63E0778C92DBDC1791E735940D0D6" type="email">saabalde@gmail.com</mods:nameIdentifier>
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<mods:namePart id="31887C67D6ECAD1E212CF0D852504E7B">Pardos-Blas, José R.</mods:namePart>
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<mods:affiliation id="A61F68F9811D4EE208A221D47D21A3B0">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain. &amp; Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain.</mods:affiliation>
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<mods:namePart id="AD8C6A7A3284958AA942F09EE0E644C3">Zardoya, Rafael</mods:namePart>
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<mods:title id="71C7A0AF72B6D9594940AFE06DE58943">European Journal of Taxonomy</mods:title>
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<mods:part id="A8BFFE10CF891A75CBE3749A1C732F4C">
<mods:date id="8D40EAA222D413F24E1530CB45D7EE77">2020</mods:date>
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<treatment id="AE38686E301EFFCFFE31F886FC540A20" ID-GBIF-Taxon="248303140" LSID="urn:lsid:plazi:treatment:AE38686E301EFFCFFE31F886FC540A20" httpUri="http://treatment.plazi.org/id/AE38686E301EFFCFFE31F886FC540A20" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72" scope_class="Gastropoda" scope_family="Conidae">
<subSubSection id="6E8B8AF3301EFFC1FE31F886FB910E24" box="[454,1134,1822,1849]" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="262ED978301EFFC1FE31F886FB910E24" blockId="71.[454,1134,1822,1883]" box="[454,1134,1822,1849]" pageId="71" pageNumber="72">
<heading id="7D666E14301EFFC1FE31F886FB910E24" box="[454,1134,1822,1849]" centered="true" fontSize="11" level="2" pageId="71" pageNumber="72" reason="2">
<taxonomicName id="E191A2FB301EFFC1FE31F886FC1A0E25" ID-CoL="7TKV4" authority="(Boyer &amp; Pelorce, 2009)" authorityName="Tenorio &amp; Abalde &amp; Pardos-Blas &amp; Cmim" authorityYear="2020" baseAuthorityName="Boyer &amp; Pelorce" baseAuthorityYear="2009" box="[454,997,1822,1849]" class="Gastropoda" family="Conidae" genus="Varioconus" kingdom="Animalia" order="Neogastropoda" pageId="71" pageNumber="72" phylum="Mollusca" rank="species" species="tacomae" status="comb. nov.">
<emphasis id="14E5056A301EFFC1FE31F886FD440E24" bold="true" box="[454,699,1822,1849]" italics="true" pageId="71" pageNumber="72">Varioconus tacomae</emphasis>
(
<bibRefCitation id="4200A489301EFFC1FD3CF886FC230E25" author="Boyer &amp; Pelorce" box="[715,988,1822,1848]" firstAuthor="Boyer" pageId="71" pageNumber="72" pagination="25 - 32" refId="ref41743" refString="Boyer F. &amp; Pelorce J. 2009. Description d'un nouveau Conus (Gastropoda: Conidae) du Senegal dans le groupe Conus mediterraneus. Novapex 10 (1): 25 - 32. Available from https: // www. biodiversitylibrary. org / page / 42270880 [accessed 29 Mar. 2020]." type="journal article" year="2009">Boyer &amp; Pelorce, 2009</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0FD6B811301EFFC1FC1BF886FB910E24" box="[1004,1134,1822,1849]" pageId="71" pageNumber="72" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6E8B8AF3301EFFC1FD1CF8D9FCB70E46" box="[747,840,1857,1883]" pageId="71" pageNumber="72" type="description">
<paragraph id="262ED978301EFFC1FD1CF8D9FCB70E46" blockId="71.[454,1134,1822,1883]" box="[747,840,1857,1883]" pageId="71" pageNumber="72">
<figureCitation id="BEAAC5FD301EFFC1FD1CF8D9FCB70E46" box="[747,840,1857,1883]" captionStart="Fig" captionStartId="72.[189,232,1610,1636]" captionTargetBox="[189,1397,265,1580]" captionTargetId="figure@72.[189,1398,265,1583]" captionTargetPageId="72" captionText="Fig. 11. A. Conus mercator Linnaeus, 1758, lectotype, 24.0 mm (LSL 254). B. Varioconus mercator (Linnaeus, 1758) comb. nov., 34.7 mm (MNCN 15.05/78482). C. Varioconus mercator f. stimpsonorum (Cossignani &amp; Allary, 2019), 39.1 mm (MNCN 15.05/78419). D. Lautoconus fernandi Petuch &amp; Berschauer, 2018, holotype, 35.4 mm (MNHN IM-2000-34012). E. Lautoconus gambiensis Petuch &amp; Berschauer, 2018, holotype, 21.6 mm (MNHN IM-2000-34013). F. Lautoconus rikae Petuch &amp; Berschauer, 2018, holotype, 17.9 mm (MNHN IM-2000-34014). G. Conus orri (Ninomiya &amp; da Motta, 1982), paratype, 30.1 mm (SNMS ZI0091234). H. Conus reticulatus Born, 1778, lectotype, 25 mm (NHMW Mollusca 14159). I. Varioconus reticulatus (Born, 1778) comb. nov., 29.5 mm (MNCN 15.05/78439). J. Conus tacomae Boyer &amp; Pelorce, 2009, holotype, 17 mm (MNHN IM-2000- 21046). K. Conus trencarti Nolf &amp; Verstraeten, 2008, holotype, 26.3 mm (MNHN IM-2000-21455). L. Varioconus unifasciatus (Kiener, 1850) comb. nov., 39.8 mm (MNCN 15.05/78451). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3890462" httpUri="https://zenodo.org/record/3890462/files/figure.png" pageId="71" pageNumber="72">Fig. 11J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6E8B8AF3301EFFC1FF4AF81BFCC70E80" box="[189,824,1923,1949]" pageId="71" pageNumber="72" type="reference_group">
<paragraph id="262ED978301EFFC1FF4AF81BFCC70E80" blockId="71.[189,824,1923,1949]" box="[189,824,1923,1949]" pageId="71" pageNumber="72">
<treatmentCitationGroup id="0681FE56301EFFC1FF4AF81BFCC70E80" box="[189,824,1923,1949]" pageId="71" pageNumber="72">
<treatmentCitation id="A730FF69301EFFC1FF4AF81BFD470E80" author="Boyer F. &amp; Pelorce J." box="[189,696,1923,1949]" page="26" pageId="71" pageNumber="72" year="2009">
<taxonomicName id="E191A2FB301EFFC1FF4AF81BFD470E80" ID-CoL="XYDN" authority="Boyer &amp; Pelorce, 2009: 26" authorityName="Boyer &amp; Pelorce" authorityPageNumber="26" authorityYear="2009" box="[189,696,1923,1949]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="71" pageNumber="72" phylum="Mollusca" rank="species" species="tacomae">
<emphasis id="14E5056A301EFFC1FF4AF81BFE8D0E80" box="[189,370,1923,1949]" italics="true" pageId="71" pageNumber="72">Conus tacomae</emphasis>
<bibRefCitation id="4200A489301EFFC1FE8DF81BFD470E80" author="Boyer &amp; Pelorce" box="[378,696,1923,1949]" firstAuthor="Boyer" page="26" pageId="71" pageNumber="72" pagination="25 - 32" refId="ref41743" refString="Boyer F. &amp; Pelorce J. 2009. Description d'un nouveau Conus (Gastropoda: Conidae) du Senegal dans le groupe Conus mediterraneus. Novapex 10 (1): 25 - 32. Available from https: // www. biodiversitylibrary. org / page / 42270880 [accessed 29 Mar. 2020]." type="journal article" year="2009">Boyer &amp; Pelorce, 2009: 26</bibRefCitation>
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, figs 9–12.
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</paragraph>
</subSubSection>
<caption id="72EE89F03011FFCEFF4AF9D2FEE80EF5" ID-DOI="http://doi.org/10.5281/zenodo.3890462" ID-Zenodo-Dep="3890462" httpUri="https://zenodo.org/record/3890462/files/figure.png" pageId="72" pageNumber="73" startId="72.[189,232,1610,1636]" targetBox="[189,1397,265,1580]" targetPageId="72">
<paragraph id="262ED9783011FFCEFF4AF9D2FEE80EF5" blockId="72.[189,1399,1610,2024]" pageId="72" pageNumber="73">
<emphasis id="14E5056A3011FFCEFF4AF9D2FEC60F79" bold="true" box="[189,313,1610,1636]" pageId="72" pageNumber="73">Fig. 11. A</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFEBDF9D3FD330F78" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[330,716,1610,1637]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="mercator">
<emphasis id="14E5056A3011FFCEFEBDF9D3FDF60F79" box="[330,521,1611,1637]" italics="true" pageId="72" pageNumber="73">Conus mercator</emphasis>
<bibRefCitation id="4200A4893011FFCEFDE4F9D3FD330F78" author="Linnaeus" box="[531,716,1610,1637]" firstAuthor="Linnaeus" pageId="72" pageNumber="73" refId="ref44943" refString="Linnaeus C. 1758. Systema Naturae per regna tria naturae, secundum classes ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio Decima, Reformata. Laurentii Salvii, Stockholm [Holmiae]. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
, lectotype, 24.0 mm (LSL 254). 
<emphasis id="14E5056A3011FFCEFBADF9D3FB910F78" bold="true" box="[1114,1134,1611,1637]" pageId="72" pageNumber="73">B</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFB89F9D3FE7B0F94" authority="(Linnaeus, 1758)" authorityName="Tenorio &amp; Abalde &amp; Pardos-Blas &amp; Cmim" authorityYear="2020" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Conidae" genus="Varioconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="mercator" status="comb. nov.">
<emphasis id="14E5056A3011FFCEFB89F9D3FA890F79" box="[1150,1398,1611,1636]" italics="true" pageId="72" pageNumber="73">Varioconus mercator</emphasis>
(
<bibRefCitation id="4200A4893011FFCEFF32F9F6FE830F94" author="Linnaeus" box="[197,380,1646,1673]" firstAuthor="Linnaeus" pageId="72" pageNumber="73" refId="ref44943" refString="Linnaeus C. 1758. Systema Naturae per regna tria naturae, secundum classes ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio Decima, Reformata. Laurentii Salvii, Stockholm [Holmiae]. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0FD6B8113011FFCEFE7CF9F6FDF10F95" box="[395,526,1646,1672]" pageId="72" pageNumber="73" rank="species">comb. nov.</taxonomicNameLabel>
, 34.7 mm (MNCN 15.05/78482). 
<emphasis id="14E5056A3011FFCEFC6DF9F6FC500F95" bold="true" box="[922,943,1646,1672]" pageId="72" pageNumber="73">C</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFC4AF9F7FB4D0F95" authorityName="Tenorio &amp; Abalde &amp; Pardos-Blas &amp; Cmim" authorityYear="2020" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[957,1202,1647,1672]" class="Gastropoda" family="Conidae" genus="Varioconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="mercator">
<emphasis id="14E5056A3011FFCEFC4AF9F7FB4D0F95" box="[957,1202,1647,1672]" italics="true" pageId="72" pageNumber="73">Varioconus mercator</emphasis>
</taxonomicName>
f. 
<taxonomicName id="E191A2FB3011FFCEFB27F9F7FDE00FB6" authority="(Cossignani &amp; Allary, 2019)" baseAuthorityName="Cossignani &amp; Allary" baseAuthorityYear="2019" class="Gastropoda" family="Conidae" genus="Lautoconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="stimpsonorum">
<emphasis id="14E5056A3011FFCEFB27F9F7FA890F95" box="[1232,1398,1647,1672]" italics="true" pageId="72" pageNumber="73">stimpsonorum</emphasis>
(
<bibRefCitation id="4200A4893011FFCEFF32F909FDE80FB6" author="Cossignani &amp; Allary" box="[197,535,1681,1707]" firstAuthor="Cossignani" pageId="72" pageNumber="73" pagination="21 - 23" refId="ref42440" refString="Cossignani T. &amp; Allary A. 2019. Lautoconus (Lautoconus) stimpsonorum: nuova specie dal Senegal. Malacologia Mostra Mondiale 104: 21 - 23." type="journal article" year="2019">Cossignani &amp; Allary, 2019</bibRefCitation>
)
</taxonomicName>
, 39.1 mm (MNCN 15.05/78419). 
<emphasis id="14E5056A3011FFCEFC27F909FC1A0FB6" bold="true" box="[976,997,1681,1707]" pageId="72" pageNumber="73">D</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFC0DF90AFE710FD3" authority="Petuch &amp; Berschauer, 2018" authorityName="Petuch &amp; Berschauer" authorityYear="2018" class="Gastropoda" family="Conidae" genus="Lautoconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="fernandi">
<emphasis id="14E5056A3011FFCEFC0DF90AFB0B0FB6" box="[1018,1268,1681,1707]" italics="true" pageId="72" pageNumber="73">Lautoconus fernandi</emphasis>
<bibRefCitation id="4200A4893011FFCEFAF5F909FE710FD3" author="Petuch &amp; Berschauer" firstAuthor="Petuch" pageId="72" pageNumber="73" pagination="164 - 172" refId="ref45956" refString="Petuch E. J. &amp; Berschauer D. P. 2018. A new Lautoconus species radiation from Gambia, West Africa. The Festivus 50 (3): 164 - 172." type="journal article" year="2018">Petuch &amp; Berschauer, 2018</bibRefCitation>
</taxonomicName>
, holotype, 35.4 mm (MNHN IM-2000-34012). 
<emphasis id="14E5056A3011FFCEFC32F92CFC260FD3" bold="true" box="[965,985,1716,1742]" pageId="72" pageNumber="73">E</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFC1EF92DFE6B0FEF" authority="Petuch &amp; Berschauer, 2018" authorityName="Petuch &amp; Berschauer" authorityYear="2018" class="Gastropoda" family="Conidae" genus="Lautoconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="gambiensis">
<emphasis id="14E5056A3011FFCEFC1EF92DFB010FD3" box="[1001,1278,1716,1742]" italics="true" pageId="72" pageNumber="73">Lautoconus gambiensis</emphasis>
<bibRefCitation id="4200A4893011FFCEFAF0F92CFE6B0FEF" author="Petuch &amp; Berschauer" firstAuthor="Petuch" pageId="72" pageNumber="73" pagination="164 - 172" refId="ref45956" refString="Petuch E. J. &amp; Berschauer D. P. 2018. A new Lautoconus species radiation from Gambia, West Africa. The Festivus 50 (3): 164 - 172." type="journal article" year="2018">Petuch &amp; Berschauer, 2018</bibRefCitation>
</taxonomicName>
, holotype, 21.6 mm (MNHN IM-2000-34013). 
<emphasis id="14E5056A3011FFCEFC03F940FBF90FEF" bold="true" box="[1012,1030,1752,1778]" pageId="72" pageNumber="73">F</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFBEBF940FE6C0E08" authority="Petuch &amp; Berschauer, 2018" authorityName="Petuch &amp; Berschauer" authorityYear="2018" class="Gastropoda" family="Conidae" genus="Lautoconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="rikae">
<emphasis id="14E5056A3011FFCEFBEBF940FB0E0FEC" box="[1052,1265,1751,1777]" italics="true" pageId="72" pageNumber="73">Lautoconus rikae</emphasis>
<bibRefCitation id="4200A4893011FFCEFAF7F94FFE6C0E08" author="Petuch &amp; Berschauer" firstAuthor="Petuch" pageId="72" pageNumber="73" pagination="164 - 172" refId="ref45956" refString="Petuch E. J. &amp; Berschauer D. P. 2018. A new Lautoconus species radiation from Gambia, West Africa. The Festivus 50 (3): 164 - 172." type="journal article" year="2018">Petuch &amp; Berschauer, 2018</bibRefCitation>
</taxonomicName>
, holotype, 17.9 mm (MNHN IM-2000-34014). 
<emphasis id="14E5056A3011FFCEFC1EF962FBFF0E09" bold="true" box="[1001,1024,1786,1812]" pageId="72" pageNumber="73">G</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFBE2F963FEA20E25" authority="(Ninomiya &amp; da Motta, 1982)" baseAuthorityName="Ninomiya &amp; da Motta" baseAuthorityYear="1982" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="orri">
<emphasis id="14E5056A3011FFCEFBE2F963FB630E09" box="[1045,1180,1787,1813]" italics="true" pageId="72" pageNumber="73">Conus orri</emphasis>
(Ninomiya &amp; 
<bibRefCitation id="4200A4893011FFCEFAACF963FEAA0E25" author="da Motta" firstAuthor="da Motta" pageId="72" pageNumber="73" pagination="1 - 20" refId="ref43066" refString="da Motta A. J. 1982. Seventeen new cone shell names (Gastropoda: Conidae). Publicacoes Ocasionais da Sociedade Portuguesa de Malacologia 1: 1 - 20." type="journal article" year="1982">da Motta, 1982</bibRefCitation>
)
</taxonomicName>
, paratype, 30.1 mm (SNMS ZI0091234). 
<emphasis id="14E5056A3011FFCEFC96F886FC870E25" bold="true" box="[865,888,1822,1848]" pageId="72" pageNumber="73">H</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFC7DF886FB0C0E25" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[906,1267,1822,1848]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="reticulatus">
<emphasis id="14E5056A3011FFCEFC7DF886FBA20E25" box="[906,1117,1822,1848]" italics="true" pageId="72" pageNumber="73">Conus reticulatus</emphasis>
<bibRefCitation id="4200A4893011FFCEFB9EF886FB0C0E25" author="Born" box="[1129,1267,1822,1848]" firstAuthor="Born" pageId="72" pageNumber="73" refId="ref41674" refString="Born I. von 1778. Index Rerum Naturalium Musei Caesarei Vindobonensis. Pars Prima, Testacea. Available from http: // biology. burke. washington. edu / conus / catalogue / sources / Born, % 20 I., % 20 1778. % 20 % 20 Index % 20 Rerum % 20 Naturalium. pdf [accessed 29 Mar. 2020]." type="book" year="1778">Born, 1778</bibRefCitation>
</taxonomicName>
, lectotype, 25 mm (NHMW Mollusca 14159). 
<emphasis id="14E5056A3011FFCEFD77F8D9FD740E46" bold="true" box="[640,651,1857,1883]" pageId="72" pageNumber="73">I</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFD56F8DAFBA40E46" authority="(Born, 1778)" authorityName="Tenorio &amp; Abalde &amp; Pardos-Blas &amp; Cmim" authorityYear="2020" baseAuthorityName="Born" baseAuthorityYear="1778" box="[673,1115,1857,1883]" class="Gastropoda" family="Conidae" genus="Varioconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="reticulatus" status="comb. nov.">
<emphasis id="14E5056A3011FFCEFD56F8DAFC500E46" box="[673,943,1857,1883]" italics="true" pageId="72" pageNumber="73">Varioconus reticulatus</emphasis>
(
<bibRefCitation id="4200A4893011FFCEFC31F8D9FBAC0E46" author="Born" box="[966,1107,1857,1883]" firstAuthor="Born" pageId="72" pageNumber="73" refId="ref41674" refString="Born I. von 1778. Index Rerum Naturalium Musei Caesarei Vindobonensis. Pars Prima, Testacea. Available from http: // biology. burke. washington. edu / conus / catalogue / sources / Born, % 20 I., % 20 1778. % 20 % 20 Index % 20 Rerum % 20 Naturalium. pdf [accessed 29 Mar. 2020]." type="book" year="1778">Born, 1778</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0FD6B8113011FFCEFB9DF8D9FB0A0E41" box="[1130,1269,1857,1884]" pageId="72" pageNumber="73" rank="species">comb. nov.</taxonomicNameLabel>
, 29.5 mm (MNCN 15.05/78439). 
<emphasis id="14E5056A3011FFCEFE25F8FCFE1E0E63" bold="true" box="[466,481,1892,1918]" pageId="72" pageNumber="73">J</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFE07F8FDFC400E63" authority="Boyer &amp; Pelorce, 2009" authorityName="Boyer &amp; Pelorce" authorityYear="2009" box="[496,959,1892,1919]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="tacomae">
<emphasis id="14E5056A3011FFCEFE07F8FDFD5A0E63" box="[496,677,1893,1919]" italics="true" pageId="72" pageNumber="73">Conus tacomae</emphasis>
<bibRefCitation id="4200A4893011FFCEFD5AF8FDFC400E63" author="Boyer &amp; Pelorce" box="[685,959,1892,1919]" firstAuthor="Boyer" pageId="72" pageNumber="73" pagination="25 - 32" refId="ref41743" refString="Boyer F. &amp; Pelorce J. 2009. Description d'un nouveau Conus (Gastropoda: Conidae) du Senegal dans le groupe Conus mediterraneus. Novapex 10 (1): 25 - 32. Available from https: // www. biodiversitylibrary. org / page / 42270880 [accessed 29 Mar. 2020]." type="journal article" year="2009">Boyer &amp; Pelorce, 2009</bibRefCitation>
</taxonomicName>
, holotype, 17 mm (MNHN IM-2000- 21046). 
<emphasis id="14E5056A3011FFCEFED6F810FEC70EBF" bold="true" box="[289,312,1928,1954]" pageId="72" pageNumber="73">K</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFEBDF810FCBE0EBF" authority="Nolf &amp; Verstraeten, 2008" authorityName="Nolf &amp; Verstraeten" authorityYear="2008" box="[330,833,1927,1954]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="trencarti">
<emphasis id="14E5056A3011FFCEFEBDF810FDFA0EBC" box="[330,517,1928,1954]" italics="true" pageId="72" pageNumber="73">Conus trencarti</emphasis>
<bibRefCitation id="4200A4893011FFCEFDF9F81FFCBE0EBF" author="Nolf &amp; Verstraeten" box="[526,833,1927,1954]" firstAuthor="Nolf" pageId="72" pageNumber="73" pagination="1 - 3" refId="ref45549" refString="Nolf F. &amp; Verstraeten J. 2008. Conus trencarti (Mollusca: Gastropoda: Conidae): a new cone from Senegal. Neptunea 7 (4): 1 - 3." type="journal article" year="2008">Nolf &amp; Verstraeten, 2008</bibRefCitation>
</taxonomicName>
, holotype, 26.3 mm (MNHN IM-2000-21455). 
<emphasis id="14E5056A3011FFCEFF4AF833FF2E0ED8" bold="true" box="[189,209,1963,1989]" pageId="72" pageNumber="73">L</emphasis>
. 
<taxonomicName id="E191A2FB3011FFCEFF15F833FD4A0ED8" authority="(Kiener, 1850)" authorityName="Tenorio &amp; Abalde &amp; Pardos-Blas &amp; Cmim" authorityYear="2020" baseAuthorityName="Kiener" baseAuthorityYear="1850" box="[226,693,1962,1989]" class="Gastropoda" family="Conidae" genus="Varioconus" kingdom="Animalia" order="Neogastropoda" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="unifasciatus" status="comb. nov.">
<emphasis id="14E5056A3011FFCEFF15F833FE020ED8" box="[226,509,1963,1989]" italics="true" pageId="72" pageNumber="73">Varioconus unifasciatus</emphasis>
(Kiener, 1850)
</taxonomicName>
<taxonomicNameLabel id="0FD6B8113011FFCEFD48F833FCBA0ED8" box="[703,837,1963,1989]" pageId="72" pageNumber="73" rank="species">comb. nov.</taxonomicNameLabel>
, 39.8 mm (MNCN 15.05/78451). Scale bars = 10 mm.
</paragraph>
</caption>
<subSubSection id="6E8B8AF33010FFCFFF4AFE93FECA0887" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="262ED9783010FFCFFF4AFE93FE550838" blockId="73.[189,426,267,293]" box="[189,426,267,293]" pageId="73" pageNumber="74">
<heading id="7D666E143010FFCFFF4AFE93FE550838" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="14E5056A3010FFCFFF4AFE93FE550838" bold="true" box="[189,426,267,293]" pageId="73" pageNumber="74">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="262ED9783010FFCFFF1DFEA2FEA30849" blockId="73.[189,1399,314,410]" box="[234,348,314,340]" pageId="73" pageNumber="74">
<emphasis id="14E5056A3010FFCFFF1DFEA2FEA30849" bold="true" box="[234,348,314,340]" pageId="73" pageNumber="74">
<typeStatus id="F92A67DA3010FFCFFF1DFEA2FEA30849" box="[234,348,314,340]" pageId="73" pageNumber="74" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="262ED9783010FFCFFF4AFEC5FECA0887" blockId="73.[189,1399,314,410]" pageId="73" pageNumber="74">
<materialsCitation id="96F9D3253010FFCFFF4AFEC5FECA0887" ID-GBIF-Occurrence="2634681329" collectionCode="MNHN" country="Senegal" elevation="10" location="East of Goree Island" municipality="Tacoma'" pageId="73" pageNumber="74" specimenCode="MNHN IM-2000-21046" specimenCount="1" typeStatus="holotype">
<collectingCountry id="5E8699E83010FFCFFF4AFEC5FEBD086A" box="[189,322,349,375]" name="Senegal" pageId="73" pageNumber="74">SENEGAL</collectingCountry>
• 
<quantity id="E169749D3010FFCFFEA8FEC5FE48086A" box="[351,439,349,375]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="73" pageNumber="74" unit="mm" value="17.0">17 mm</quantity>
; 
<location id="234E8FA33010FFCFFE31FEC5FD3E086A" LSID="urn:lsid:plazi:treatment:AE38686E301EFFCFFE31F886FC540A20:234E8FA33010FFCFFE31FEC5FD3E086A" box="[454,705,349,375]" country="Senegal" municipality="Tacoma'" name="East of Goree Island" pageId="73" pageNumber="74">East of Gorée Island</location>
, ‘ 
<collectingMunicipality id="C64A43023010FFCFFD24FEC5FCC3086A" box="[723,828,349,375]" pageId="73" pageNumber="74">Tacoma’</collectingMunicipality>
wreck; 
<elevation id="ADBC3E4B3010FFCFFC57FEC5FBB5086A" box="[928,1098,349,375]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="73" pageNumber="74" unit="m" value="10.0" valueMax="13.0" valueMin="7.0">
<quantity id="E169749D3010FFCFFC57FEC5FC04086A" box="[928,1019,349,375]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="73" pageNumber="74" unit="m" value="10.0" valueMax="13.0" valueMin="7.0">7–13 m</quantity>
depth
</elevation>
; 
<specimenCode id="763771033010FFCFFBAFFEC5FA88086B" box="[1112,1399,348,375]" pageId="73" pageNumber="74">MNHN IM-2000-21046</specimenCode>
(
<figureCitation id="BEAAC5FD3010FFCFFF31FE18FED70887" box="[198,296,384,410]" captionStart="Fig" captionStartId="72.[189,232,1610,1636]" captionTargetBox="[189,1397,265,1580]" captionTargetId="figure@72.[189,1398,265,1583]" captionTargetPageId="72" captionText="Fig. 11. A. Conus mercator Linnaeus, 1758, lectotype, 24.0 mm (LSL 254). B. Varioconus mercator (Linnaeus, 1758) comb. nov., 34.7 mm (MNCN 15.05/78482). C. Varioconus mercator f. stimpsonorum (Cossignani &amp; Allary, 2019), 39.1 mm (MNCN 15.05/78419). D. Lautoconus fernandi Petuch &amp; Berschauer, 2018, holotype, 35.4 mm (MNHN IM-2000-34012). E. Lautoconus gambiensis Petuch &amp; Berschauer, 2018, holotype, 21.6 mm (MNHN IM-2000-34013). F. Lautoconus rikae Petuch &amp; Berschauer, 2018, holotype, 17.9 mm (MNHN IM-2000-34014). G. Conus orri (Ninomiya &amp; da Motta, 1982), paratype, 30.1 mm (SNMS ZI0091234). H. Conus reticulatus Born, 1778, lectotype, 25 mm (NHMW Mollusca 14159). I. Varioconus reticulatus (Born, 1778) comb. nov., 29.5 mm (MNCN 15.05/78439). J. Conus tacomae Boyer &amp; Pelorce, 2009, holotype, 17 mm (MNHN IM-2000- 21046). K. Conus trencarti Nolf &amp; Verstraeten, 2008, holotype, 26.3 mm (MNHN IM-2000-21455). L. Varioconus unifasciatus (Kiener, 1850) comb. nov., 39.8 mm (MNCN 15.05/78451). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3890462" httpUri="https://zenodo.org/record/3890462/files/figure.png" pageId="73" pageNumber="74">Fig. 11J</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6E8B8AF33010FFCFFF4AFE21FE4F0B1F" pageId="73" pageNumber="74" type="distribution">
<paragraph id="262ED9783010FFCFFF4AFE21FDFD08CE" blockId="73.[189,514,441,467]" box="[189,514,441,467]" pageId="73" pageNumber="74">
<heading id="7D666E143010FFCFFF4AFE21FDFD08CE" bold="true" box="[189,514,441,467]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="14E5056A3010FFCFFF4AFE21FDFD08CE" bold="true" box="[189,514,441,467]" pageId="73" pageNumber="74">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="262ED9783010FFCFFF4AFE70FE4F0B1F" blockId="73.[189,432,488,515]" box="[189,432,488,515]" pageId="73" pageNumber="74">
<collectingRegion id="E455179A3010FFCFFF4AFE70FEFA0B1F" box="[189,261,488,514]" country="Senegal" name="Dakar" pageId="73" pageNumber="74">Dakar</collectingRegion>
area, 
<collectingCountry id="5E8699E83010FFCFFEBCFE70FE530B1F" box="[331,428,488,514]" name="Senegal" pageId="73" pageNumber="74">Senegal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6E8B8AF33010FFCFFF4AFDBAFEBA0B76" pageId="73" pageNumber="74" type="conservation">
<paragraph id="262ED9783010FFCFFF4AFDBAFD650B21" blockId="73.[189,666,546,572]" box="[189,666,546,572]" pageId="73" pageNumber="74">
<heading id="7D666E143010FFCFFF4AFDBAFD650B21" bold="true" box="[189,666,546,572]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="14E5056A3010FFCFFF4AFDBAFD650B21" bold="true" box="[189,666,546,572]" pageId="73" pageNumber="74">Conservation status in IUCN Red List</emphasis>
</heading>
</paragraph>
<paragraph id="262ED9783010FFCFFF4AFDC9FEBA0B76" blockId="73.[189,325,593,619]" box="[189,325,593,619]" pageId="73" pageNumber="74">Vulnerable.</paragraph>
</subSubSection>
<subSubSection id="6E8B8AF33010FFCFFF4AFD13FC540A20" pageId="73" pageNumber="74" type="discussion">
<paragraph id="262ED9783010FFCFFF4AFD13FED00BB8" blockId="73.[189,303,651,677]" box="[189,303,651,677]" pageId="73" pageNumber="74">
<heading id="7D666E143010FFCFFF4AFD13FED00BB8" bold="true" box="[189,303,651,677]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="14E5056A3010FFCFFF4AFD13FED00BB8" bold="true" box="[189,303,651,677]" pageId="73" pageNumber="74">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="262ED9783010FFCFFF4AFD21FC540A20" blockId="73.[189,1399,697,829]" pageId="73" pageNumber="74">
This species has not been molecularly examined. In the original description it was compared to 
<taxonomicName id="E191A2FB3010FFCFFADCFD22FEC80BEB" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="desidiosus">
<emphasis id="14E5056A3010FFCFFADCFD22FEC80BEB" italics="true" pageId="73" pageNumber="74">Conus desidiosus</emphasis>
</taxonomicName>
(which does not live in 
<collectingCountry id="5E8699E83010FFCFFDB4FD44FD5B0BEA" box="[579,676,732,759]" name="Senegal" pageId="73" pageNumber="74">Senegal</collectingCountry>
), 
<taxonomicName id="E191A2FB3010FFCFFD42FD44FCC40BEB" authorityName="Tenorio &amp; Abalde &amp; Pardos-Blas &amp; Cmim" authorityYear="2020" baseAuthorityName="Kiener" baseAuthorityYear="1846" box="[693,827,732,758]" class="Gastropoda" family="Conidae" genus="Varioconus" kingdom="Animalia" order="Neogastropoda" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="bruguieresi">
<emphasis id="14E5056A3010FFCFFD42FD44FCC40BEB" box="[693,827,732,758]" italics="true" pageId="73" pageNumber="74">bruguieresi</emphasis>
</taxonomicName>
and 
<taxonomicName id="E191A2FB3010FFCFFC98FD44FBFA0BEB" authorityName="Petuch" authorityYear="1975" box="[879,1029,732,758]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="echinophilus">
<emphasis id="14E5056A3010FFCFFC98FD44FBFA0BEB" box="[879,1029,732,758]" italics="true" pageId="73" pageNumber="74">echinophilus</emphasis>
</taxonomicName>
. The species is morphologically related to the latter two, and might represent a variation of one of these. We consider this species as provisionally valid only, pending future molecular examination.
</paragraph>
</subSubSection>
</treatment>
</document>