Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan Author Shimizu, So Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan & DC and Overseas Challenge Program for Young Researchers, Japan Society for the Promotion of Science, Tokyo, Japan & Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK https://orcid.org/0000-0002-5202-4552 parasitoidwasp.sou@gmail.com Author Broad, Gavin R. Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK https://orcid.org/0000-0001-7223-5333 Author Maeto, Kaoru Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan text ZooKeys 2020 990 1 144 http://dx.doi.org/10.3897/zookeys.990.55542 journal article http://dx.doi.org/10.3897/zookeys.990.55542 1313-2970-990-1 7B73642C278D40F89091B26213C9A704 9F51F78CD53F5005A305DE65494002C4 Enicospilus tripartitus Chiu, 1954 Figure 49 Enicospilus tripartitus Chiu, 1954: 36; HT ♀ from Taiwan, TARI, examined. Specimens examined. Total of 67 specimens (39♀♀26♂♂ and 2 unsexed): China (1♀), India (1♂), Japan (12♀♀14♂♂ and 1 unsexed), Nepal (24♀♀8♂♂ and 1 unsexed), Taiwan (2♀♀), unknown (3♂♂). Type series: HT ♀ of Enicospilus tripartitus Chiu, 1954, Taihoku, TAIWAN, 27.VIII.1937, J. Sonan leg. (TARI); PT ♂ of E. tripartitus , no data (NHMUK). Distribution. Eastern Palaearctic and Oriental regions ( Yu et al. 2016 ). JAPAN: [ Tohoku ] Miyagi*; [Hokuriku] Niigata* and Ishikawa*; [ Kanto-Koshin ] Kanagawa*; [ Tokai ] Shizuoka*, Aichi*, and Mie*; [ Chugoku ] Shimane* and Hiroshima ( Maeto and Shimizu 2019 ; present study); [Shikoku] Tokushima* and Kochi *; [ Kyushu ] Ōita ( Chiu 1954 ); [ Ryukyus ] Kagoshima* and Okinawa ( Chiu 1954 ; Shimizu 2020 ; present study). *New records. Bionomics. Unknown. Differential diagnosis. This species resembles E. laqueatus , E. pseudantennatus , and E. vestigator in the shapes of the sclerites, but can easily be distinguished by the dense and stout setae and punctures of the outer mandibular surface (Fig. 49B, D ), deep basal concavity of the outer mandibular surface (Fig. 49B, D ), etc., as summarised in Table 6 . Figure 49. Enicospilus tripartitus Chiu, 1954 ♂ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing.