Phylogenomic placement and revision of Iranattus Prószyński, 1992 jumping spiders (Salticidae, Plexippini, Plexippina) Author Marathe, Kiran 0000-0002-7364-3475 Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada & National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bengaluru, 560065, India Author Tripathi, Rishikesh 0000-0002-9192-4609 Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, 680 125, India Author Sudhikumar, Ambalaparambil V. https: // orcid. org / 0000 - 0002 - 4479 - 4995 Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, 680 125, India Author Maddison, Wayne P. https: // orcid. org / 0000 - 0003 - 4953 - 4575 Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada text Zoosystematics and Evolution 2024 2024-05-14 100 2 531 542 journal article 10.3897/zse.100.122034 4488FFD3-5621-439E-9253-058E974EB0B3 Iranattus Prószyński, 1992 Figs 2 – 7 , 8 – 13 , 14 – 19 , 20 – 23 , 24 – 29 , 30 – 33 , 34–41 Iranattus Prószyński, 1992: 97 – 98 , f. 35 – 40. Monomotapa Wesołowska, 2000: 159 , f. 42–46 (synonymized by Prószyński, 2017: 36 .). Type species. Iranattus rectangularis Prószyński, 1992 . Species included. Iranattus principalis ( Wesołowska, 2000 ) ; Iranattus rectangularis Prószyński, 1992 . Diagnosis. The remarkably long third legs of Iranattus (Figs 15 , 18 , 30 , 32 ) and scoop-shaped cymbial apophysis (Fig. 4 ) differentiate it from all other plexippines. The very robust carapace, bulging outward at the PLE and bearing the PLEs on tubercles, is unusual but shared also with Afrobeata Caporiacco, 1941 , and Vailimia Kammerer, 2006 . Vailimia especially might be confused with Iranattus , as they share erect hairs on the carapace (see Figs 34 , 38 , 41 ) and a compact crouch stance, but, besides the cymbial apophysis and long third legs, Iranattus also has a shorter embolus lacking membrane (membrane-accompanied long embolus in Vailimia ), a short RTA (long and curved in Vailimia ), and two distinct deep conical ECPs (absent in Vailimia ). From Afrobeata , Iranattus differs in having longer third legs, a cymbial apophysis (lacking in Afrobeata ), a shorter embolus (longer in Afrobeata ), a simple short RTA (bifurcated in Afrobeata ), shorter copulatory ducts (long in Afrobeata ), and deep conical ECPs (shallow in Afrobeata ). Some other plexippines have cymbial apophyses ( Plexippoides Prószyński, 1984 ; Epeus Peckham & Peckham, 1886 ; and Erasinus Simon, 1899 ), but their apophyses are different in shape — in Iranattus , a long, broad blade with a rounded tip, concave in front so as to form a scoop; in Plexippoides , sharply pointed, for example. Figures 2 – 7. Iranattus principalis genitalia drawings. 2. Male left palp, ventral view (DDKM 21.089); 3. Ditto, oblique view (DDKM 21.089); 4. Ditto, oblique view, closeup of the cymbial apophysis (DDKM 21.089); 5. Ditto, retrolateral view (DDKM 21.089); 6. Epigyne, ventral view (DDKM 21.090); 7. Vulva, dorsal view (DDKM 21.089). Scale bars: 0.2 mm.