Noideattella and Tolegnaro, Two New Genera of Goblin Spiders from Madagascar, with Comments on the Gamasomorphoid and Silhouettelloid Oonopids (Araneae, Oonopidae) Author Álvarez-Padilla, Fernando Author Ubick, Darrell Author Griswold, Charles E. text American Museum Novitates 2012 2012-05-25 2012 3745 1 76 http://www.bioone.org/doi/abs/10.1206/3745.2 journal article 2558 10.1206/3745.2 25e1eb0b-a2ad-4c0f-aaec-21d56c6fde5e 0003-0082 5360355 Noideattella fantara , new species Figures 231–256 , 376 TYPES: Male holotype and female allotype from Berlese and sifted leaf litter of a spiny forest/thicket at an elevation of 150 m at Parc National d’Andohahela , Forêt de Manantalinjo , 33.6 km 63° ENE Amboasary , 7.6 km 99° E Hazofotsy , 24°49′1″S 46° 36′36″E , Toliara Province , 12–16 January 2002 , Fisher-Griswold Arthropod Team , deposited in CASC ( PBI_ OON 00036162 ). The species epithet, a noun in apposition, is taken from the Malagasy word for falling star . DIAGNOSIS: N. fantara can be differentiated from other Noideattella species by the following combination of characters: habitus pale orange; carapace surface of elevated portion of pars cephalica and sides granulate ( figs. 245, 246 ); sternum surface finely reticulate, without pits and microsculpture only in furrows (figs. 235, 243, 244 ); male cheliceral paturon inner margin with brush of modified setae ( figs. 247–249 ); abdomen cylindrical, book lung covers small and round, epigastric scutum length about at half of the abdomen (figs. 232, 233). DESCRIPTION: Male: Total length 1.4. Cephalothorax: Carapace pale orange. Sternum longer than wide, pale orange, anterior margin with interrupted transverse groove. Mouthparts: Chelicerae, endites, and labium pale orange. Sternum anterior margin with interrupted transverse groove ( fig. 242 ). Abdomen: Dorsal scutum pale orange, middle surface and sides reticulate. Postepigastric scutum pale orange, long, semicircular, covering nearly full length of abdominal length. Male genital opening slightly protruded, epigastric scutum apodemes small, separated from the anterior book lung opening and not connected by a furrow. Posterior spiracles connected by a groove ( fig. 241 ). Legs: Yellow, without color pattern. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I, II p1-1-1; r1-1-1; IV p0-0-1; metatarsi: I, II p1-1-0; r1-1-0. Tarsi I to IV superior claw teeth not examined in detail. Tarsal organ with two sensilla visible. Genitalia: Epigastric region with sperm pore situated at level of anterior spiracles ( fig. 241 ). Palp proximal segments yellow; cymbium yellow, with distal patch of setae; bulb yellow (figs. 236, 238). Embolus dorsal lamella with at least two distal apophyses, ventral lamella with at least three apophyses (figs. 253–256). Palp as in figures 250–252. Female (PBI_OON 0003270): Total length 1.8. As in male except as noted. Legs: Metatarsi III and IV weak ventral scopula absent. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I, II p1-1-1; r1-1-1; IV p0-0-1; metatarsi: I, II p1-1-0; r1-1-0. Genitalia as in figures 239 and 240. Figs. 231–240. Noideattella fantara , new species . 231. Habitus male, dorsal view. 232. Same, ventral view. 233. Same, lateral view. 234. Cephalothorax, dorsal view. 235. Same, ventral view. 236. Palp male, prolateral view. 237. Cephalothorax male, anterior view. 238. Palp male, retrolateral view. 239. Genitalia female, ventral view. 240. Same, dorsal view. Scale bars = 0.2 mm , except genitalia: 0.1 mm . Figs. 241–249. Noideattella fantara , new species . 241. Epigastric region male, ventral view. 242. Endites male, ventral view. 243. Sternum male, ventral view. 244. Infracoxal grooves male, ventral view. 245. Cephalothorax male, dorsal view. 246. Same, lateral view. 247. Cephalothorax male, anterior view. 248. Chelicerae male, anterior view. 249. Chelicerae modified setae male, anterior view. Scale bars = 50 µm, except 249: 10 µm. Figs. 250–256. Noideattella fantara , new species . 250. Palp male, prolateral view. 251. Same, retrolateral view. 252. Same, dorsal view. 253. Embolus, prolateral view. 254. Same, retrolateral view. 255. Same, dorsal view. 256. Same, ventral view. Scale bars = 50 µm, except 253–256: 10 µm. MATERIAL EXAMINED: N = 33. Madagascar : Antananarivo : Reserve Speciale d’Ambohitantely , Forêt d’Ambohitantely , 24.1 km 59° NE d’Ankazobe , montane rainforest , 1620 m , S18.00277° , E47.00444° , Apr. 17–22, 2001 , Fisher-Griswold Arthropod Team, 1♂ ( CASC PBI _ OON 3980 ) . Antsiranana : Reserve Speciale d’Anjajaharibe , Sud: 12.2 km WSW Befingotra , montane rainforest , 1985 m , S14.00694° , E49.00722° , Nov. 25, 1994 , Fisher-Griswold Arthropod Team , 1♂ ( CASC PBI _ OON 3918 ) ; Reserve Speciale l’Ankarana , 13 km 192° SSW Anivorano Nord , tropical dry forest , 210 m , S12.86361° , E49.22583° , Feb. 16–21, 2001 , Fisher- Griswold Arthropod Team , 1♀ ( CASC PBI _ OON 3270 ) . Toliara : Parc National d’Andohahela , Forêt de Manantalinjo , 33.6 km 63° ENE Amboasary , 7.6 km 99° E Hazofotsy , spiny forest thicket, 150 m , S24.81694° , E46.61000° , Jan. 12–16, 2002 , Fisher-Griswold Arthropod Team , 11♂ ( CASC PBI _OON 36386) ; Parc National de Tsimanampetsotsa , Forêt de Bemanateza , 20.7 km 81° E Efoetse , 23.0 km 131° SE Beheloka , spiny forest thicket , 90 m , S23.99222° , E43.88055° , Mar. 22–26, 2002 , Fisher-Griswold Arthropod Team , 2 mixed ( CASC PBI _OON 36245) ; Reserve Speciale de Cap Sainte Marie , 12.3 km 262° W Marovato , spiny forest thicket , 200 m , S25.58166° , E45.16833° , Feb. 11–15, 2002 , Fisher-Griswold Arthropod Team , 13♂ ( CASC PBI _ OON 3342 ) ; Reserve Speciale de Cap Sainte Marie , 14.9 km 261° W Marovato , spiny forest thicket , 160 m , S25.59444° , E45.14694° , Feb. 13–19, 2002 , Fisher-Griswold Arthropod Team , 2♂ ( CASC PBI _OON 36374) .