<document id="62611551DC75D7A8F1DEC8AABD2C1ADB" ID-CLB-Dataset="29189" ID-DOI="10.11646/zootaxa.4449.1.1" ID-GBIF-Dataset="a8aa5ad0-dbcc-4e48-9211-b83b90e0aee7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1454305" ID-ZooBank="0734DC4E-2C9B-4CD2-B8FE-8F94589200E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539174719248" checkinUser="plazi" docAuthor="Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel &amp; Solodovnikov, Alexey" docDate="2018" docId="215187C84C02B363FF6BF988F315D6DA" docLanguage="en" docName="zootaxa.4449.1.1.pdf" docOrigin="Zootaxa 4449 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Laetulonthus Moore &amp; Legner 1973" docType="treatment" docVersion="7" lastPageNumber="44" masterDocId="DD68FFB04C28B348FFFCFFD4F755D564" masterDocTitle="Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae)" masterLastPageNumber="95" masterPageNumber="1" pageNumber="43" updateTime="1698571138576" updateUser="ExternalLinkService">
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<mods:title id="6B073D9F218EC89B66BD735049F3CA3A">Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="A8AD60619C962843890E4478FF851D4B">Chani-Posse, Mariana</mods:namePart>
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<mods:namePart id="B80DA137BC3D5B7B11D84039D0C3CDC5">Newton, Alfred F.</mods:namePart>
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<mods:date id="391106F5B80A203A7EB947487363CBD6">2018</mods:date>
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<subSubSection id="E1E265554C02B362FF6BF988F500D312" box="[151,597,1628,1654]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="A94736DE4C02B362FF6BF988F500D312" blockId="42.[151,597,1628,1654]" box="[151,597,1628,1654]" pageId="42" pageNumber="43">
<heading id="F20F81B24C02B362FF6BF988F500D312" bold="true" box="[151,597,1628,1654]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<taxonomicName id="6EF84D5D4C02B362FF6BF988F500D312" ID-CoL="8VRS2" authority="Moore &amp; Legner 1973" authorityName="Moore &amp; Legner" authorityYear="1973" box="[151,597,1628,1654]" class="Insecta" family="Staphylinidae" genus="Laetulonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C02B362FF6BF988F500D312" bold="true" box="[151,597,1628,1654]" pageId="42" pageNumber="43">
<emphasis id="9B8CEACC4C02B362FF6BF988F66DD312" bold="true" box="[151,312,1628,1654]" italics="true" pageId="42" pageNumber="43">Laetulonthus</emphasis>
<bibRefCitation id="CD694B2F4C02B362FEC3F988F500D312" box="[319,597,1628,1654]" pageId="42" pageNumber="43" refId="ref69624" refString="Moore, I. &amp; Legner, E. F. (1973) Laetulonthus, a new genus for Philonthus laetulus Say (Coleoptera: Staphylinidae). Journal of the New York Entomological Society, 80 (1972), 212 - 215." type="journal article">Moore &amp; Legner 1973</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<paragraph id="A94736DE4C02B362FF6BF976F55CD392" blockId="42.[151,1437,1698,1782]" pageId="42" pageNumber="43">
<treatmentCitationGroup id="89E811F04C02B362FF6BF976F55CD392" infered="true" pageId="42" pageNumber="43">
<taxonomicName id="6EF84D5D4C02B362FF6BF976F56FD3DC" ID-CoL="8VRS2" authority="Moore &amp; Legner 1973: 213" authorityName="Moore &amp; Legner" authorityPageNumber="213" authorityYear="1973" box="[151,570,1698,1720]" class="Insecta" family="Staphylinidae" genus="Laetulonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C02B362FF6BF976F64ED3DC" box="[151,283,1698,1720]" italics="true" pageId="42" pageNumber="43">Laetulonthus</emphasis>
<treatmentCitation id="285910CF4C02B362FEDDF977F56FD3DC" author="" box="[289,570,1698,1720]" page="213" pageId="42" pageNumber="43">
<bibRefCitation id="CD694B2F4C02B362FEDDF977F55DD3DC" box="[289,520,1698,1720]" pageId="42" pageNumber="43" refId="ref69624" refString="Moore, I. &amp; Legner, E. F. (1973) Laetulonthus, a new genus for Philonthus laetulus Say (Coleoptera: Staphylinidae). Journal of the New York Entomological Society, 80 (1972), 212 - 215." type="journal article">Moore &amp; Legner 1973</bibRefCitation>
: 213
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</taxonomicName>
, ex 
<taxonomicName id="6EF84D5D4C02B362FD9EF976F585D3DC" ID-CoL="8VRYR" baseAuthorityName="Tottenham" baseAuthorityYear="1955" box="[610,720,1698,1720]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C02B362FD9EF976F585D3DC" box="[610,720,1698,1720]" italics="true" pageId="42" pageNumber="43">Philonthus</emphasis>
</taxonomicName>
. 
<typeStatus id="7643887C4C02B362FD20F977F45BD3DC" box="[732,782,1699,1720]" pageId="42" pageNumber="43">Type</typeStatus>
species: 
<taxonomicName id="6EF84D5D4C02B362FC97F976F3F7D3DC" ID-CoL="4ZK46" authority="Say, 1830" authorityName="Say" authorityYear="1830" box="[875,1186,1698,1720]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="laetulus">
<emphasis id="9B8CEACC4C02B362FC97F976F363D3DC" box="[875,1078,1698,1720]" italics="true" pageId="42" pageNumber="43">Staphylinus laetulus</emphasis>
<bibRefCitation id="CD694B2F4C02B362FBC1F976F3F7D3DC" box="[1085,1186,1698,1720]" pageId="42" pageNumber="43" refId="ref71623" refString="Say, T. (1830) Descriptions of new species of North American insects, and observations on some already described. New Harmony, Indiana, pp. 1 - 41." type="book chapter">Say, 1830</bibRefCitation>
</taxonomicName>
, by original designation. Note: 
<bibRefCitation id="CD694B2F4C02B362FEFAF915F6FDD3B3" box="[262,424,1729,1751]" pageId="42" pageNumber="43" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana (1995)</bibRefCitation>
and 
<bibRefCitation id="CD694B2F4C02B362FE20F916F5D0D3B3" box="[476,645,1729,1751]" pageId="42" pageNumber="43" refId="ref67417" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article">Herman (2001d)</bibRefCitation>
give 1834 as publication date for 
<taxonomicName id="6EF84D5D4C02B362FC1BF916F304D3B3" ID-CoL="4ZK46" authorityName="Say" authorityYear="1830" box="[999,1105,1729,1751]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="laetulus">
<emphasis id="9B8CEACC4C02B362FC1BF916F304D3B3" box="[999,1105,1729,1751]" italics="true" pageId="42" pageNumber="43">S. laetulus</emphasis>
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, but its original description was actually first published in 1830.
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Note: 
<taxonomicName id="6EF84D5D4C02B362FF1DF8C9F626D252" ID-CoL="8VRS2" authorityName="Moore &amp; Legner" authorityYear="1973" box="[225,371,1821,1846]" class="Insecta" family="Staphylinidae" genus="Laetulonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C02B362FF1DF8C9F626D252" box="[225,371,1821,1846]" italics="true" pageId="42" pageNumber="43">Laetulonthus</emphasis>
</taxonomicName>
, so far as currently known, is an endemic North American genus (southernmost record is 
<collectingRegion id="6B3CF83C4C02B362FF39F895F646D23E" box="[197,275,1857,1882]" country="Mexico" name="Jalisco" pageId="42" pageNumber="43">Jalisco</collectingRegion>
) with two species at present (
<bibRefCitation id="CD694B2F4C02B362FDA2F896F426D23E" box="[606,883,1857,1882]" pageId="42" pageNumber="43" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 505–513</bibRefCitation>
).
</paragraph>
<paragraph id="A94736DE4C02B362FF6BF85DF506D2C6" blockId="42.[151,1437,1929,2026]" box="[151,595,1929,1954]" pageId="42" pageNumber="43">
<taxonomicName id="6EF84D5D4C02B362FF6BF85DF506D2C6" ID-CoL="6NT6P" authority="(Bernhauer 1910)" authorityName="Bernhauer" authorityYear="1910" baseAuthorityName="Bernhauer" baseAuthorityYear="1910" box="[151,595,1929,1954]" class="Insecta" family="Staphylinidae" genus="Laetulonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="9B8CEACC4C02B362FF6BF85DF6D3D2C6" bold="true" box="[151,390,1929,1954]" italics="true" pageId="42" pageNumber="43">Laetulonthus nobilis</emphasis>
(
<bibRefCitation id="CD694B2F4C02B362FE68F85DF51FD2C6" box="[404,586,1929,1954]" pageId="42" pageNumber="43" refId="ref61912" refString="Bernhauer, M. (1910) Beitrag zur Kenntnis der Staphyliniden-Fauna von Zentralamerika. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 60, 350 - 393." type="journal article">Bernhauer 1910</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C02B363FF6BF879F5D7D5D4" blockId="42.[151,1437,1929,2026]" lastBlockId="43.[151,1436,151,240]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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<taxonomicName id="6EF84D5D4C02B362FF6BF879F50CD2A2" ID-CoL="4GBD4" authority="Bernhauer 1910: 381" authorityName="Bernhauer" authorityPageNumber="381" authorityYear="1910" box="[151,601,1965,1990]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="9B8CEACC4C02B362FF6BF879F636D2A2" box="[151,355,1965,1990]" italics="true" pageId="42" pageNumber="43">Philonthus nobilis</emphasis>
<treatmentCitation id="285910CF4C02B362FE96F879F50CD2A2" author="" box="[362,601,1965,1990]" page="381" pageId="42" pageNumber="43">
<bibRefCitation id="CD694B2F4C02B362FE96F879F574D2A2" box="[362,545,1965,1990]" pageId="42" pageNumber="43" refId="ref61912" refString="Bernhauer, M. (1910) Beitrag zur Kenntnis der Staphyliniden-Fauna von Zentralamerika. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 60, 350 - 393." type="journal article">Bernhauer 1910</bibRefCitation>
: 381
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C02B362FD98F87AF5C9D2A2" box="[612,668,1966,1990]" pageId="42" pageNumber="43">Type</typeStatus>
locality: Porvenir, 
<collectingRegion id="6B3CF83C4C02B362FC8EF87AF352D2A2" box="[882,1031,1965,1990]" country="United States of America" name="New Mexico" pageId="42" pageNumber="43">
New 
<collectingRegion id="6B3CF83C4C02B362FC52F879F352D2A2" box="[942,1031,1965,1990]" country="Mexico" name="Mexico" pageId="42" pageNumber="43">
<collectingCountry id="D1EF764E4C02B362FC52F879F352D2A2" box="[942,1031,1965,1990]" name="Mexico" pageId="42" pageNumber="43">Mexico</collectingCountry>
</collectingRegion>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C02B362FBEDF87AF318D2A2" box="[1041,1101,1966,1990]" name="United States of America" pageId="42" pageNumber="43">USA</collectingCountry>
. 
<typeStatus id="7643887C4C02B362FBABF87AF3DAD2A2" box="[1111,1167,1966,1990]" pageId="42" pageNumber="43">Type</typeStatus>
depository: FMNH HT. Distribution: 
<collectingCountry id="D1EF764E4C02B362FEA7F806F6C3D28E" box="[347,406,2002,2026]" name="United States of America" pageId="42" pageNumber="43">USA</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C02B362FE5DF805F6AFD28E" box="[417,506,2001,2026]" country="Mexico" name="Mexico" pageId="42" pageNumber="43">
<collectingCountry id="D1EF764E4C02B362FE5DF805F6AFD28E" box="[417,506,2001,2026]" name="Mexico" pageId="42" pageNumber="43">Mexico</collectingCountry>
</collectingRegion>
. Note: distribution, also within 
<collectingRegion id="6B3CF83C4C02B362FCA0F805F4E4D28E" box="[860,945,2001,2026]" country="Mexico" name="Mexico" pageId="42" pageNumber="43">
<collectingCountry id="D1EF764E4C02B362FCA0F805F4E4D28E" box="[860,945,2001,2026]" name="Mexico" pageId="42" pageNumber="43">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C02B362FC43F805F241D28E" box="[959,1300,2001,2026]" pageId="42" pageNumber="43" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C02B362FB65F806F384D28E" box="[1177,1233,2001,2026]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2002
</bibRefCitation>
). 
<taxonomicName id="6EF84D5D4C03B363FF6BFF43F5D7D5D4" ID-CoL="8XMBD" authority="Chani-Posse &amp; Asenjo 2013" authorityName="Chani-Posse &amp; Asenjo" authorityYear="2013" box="[151,642,151,176]" class="Insecta" family="Staphylinidae" genus="Leptopeltoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C03B363FF6BFF43F668D5D4" bold="true" box="[151,317,151,176]" italics="true" pageId="43" pageNumber="44">Leptopeltoides</emphasis>
<bibRefCitation id="CD694B2F4C03B363FEB8FF43F5D7D5D4" box="[324,642,151,176]" pageId="43" pageNumber="44" refId="ref64481" refString="Chani-Posse, M. &amp; Asenjo, A. (2013) Systematics and phylogeny of the Andean genera Leptopeltus Bernhauer and Leptopeltoides gen. nov. (Coleoptera: Staphylinidae), with biogeographical notes. Zoologischer Anzeiger, 252, 440 - 456. https: // doi. org / 10.1016 / j. jcz. 2013.02.001" type="journal article">Chani-Posse &amp; Asenjo 2013</bibRefCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C03B363FF6BFF68F6CDD594" blockId="43.[151,1436,151,240]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="89E811F04C03B363FF6BFF68F6CDD594" pageId="43" pageNumber="44">
<taxonomicName id="6EF84D5D4C03B363FF6BFF68F5DED5B6" ID-CoL="8XMBD" authority="Chani-Posse &amp; Asenjo 2013: 450" authorityName="Chani-Posse &amp; Asenjo" authorityPageNumber="450" authorityYear="2013" box="[151,651,188,210]" class="Insecta" family="Staphylinidae" genus="Leptopeltoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C03B363FF6BFF68F679D5B6" box="[151,300,188,210]" italics="true" pageId="43" pageNumber="44">Leptopeltoides</emphasis>
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species: 
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<emphasis id="9B8CEACC4C03B363FCD8FF68F312D5B6" box="[804,1095,188,210]" italics="true" pageId="43" pageNumber="44">Leptopeltoides ecuatoriensis</emphasis>
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, by original designation.
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Note: 
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<emphasis id="9B8CEACC4C03B363FF22FEC9F6D6D452" box="[222,387,285,310]" italics="true" pageId="43" pageNumber="44">Leptopeltoides</emphasis>
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is a Neotropical genus with four species currently known from the Andes of Colombia and Ecuador (
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).
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<emphasis id="9B8CEACC4C03B363FF6BFE5CF68DD4C5" bold="true" box="[151,472,392,417]" italics="true" pageId="43" pageNumber="44">Leptopeltoides chingazensis</emphasis>
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<emphasis id="9B8CEACC4C03B363FF6BFE79F686D4A2" box="[151,467,429,454]" italics="true" pageId="43" pageNumber="44">Leptopeltoides chingazensis</emphasis>
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<taxonomicName id="6EF84D5D4C03B363FF6BFDCCF475D755" ID-CoL="3TSPS" authority="Chani-Posse &amp; Asenjo 2013" authorityName="Chani-Posse &amp; Asenjo" authorityYear="2013" box="[151,800,536,562]" class="Insecta" family="Staphylinidae" genus="Leptopeltoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="columbiensis">
<emphasis id="9B8CEACC4C03B363FF6BFDCCF68ED755" bold="true" box="[151,475,536,561]" italics="true" pageId="43" pageNumber="44">Leptopeltoides columbiensis</emphasis>
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<taxonomicName id="6EF84D5D4C03B363FF6BFD7DF477D7A6" ID-CoL="3TSPT" authority="Chani-Posse &amp; Asenjo 2013" authorityName="Chani-Posse &amp; Asenjo" authorityYear="2013" box="[151,802,681,706]" class="Insecta" family="Staphylinidae" genus="Leptopeltoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ecuatoriensis">
<emphasis id="9B8CEACC4C03B363FF6BFD7DF689D7A6" bold="true" box="[151,476,681,706]" italics="true" pageId="43" pageNumber="44">Leptopeltoides ecuatoriensis</emphasis>
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