Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera
Author
Winterton, Shaun L.
Author
Martins, Caleb Califre
Author
Makarkin, Vladimir
Author
Ardila-Camacho, Adrian
Author
Wang, Yongjie
text
Zootaxa
2019
2019-04-09
4581
1
1
99
journal article
27348
10.11646/zootaxa.4581.1.1
5b259853-01f6-4307-8bef-58c98ec73bd3
1175-5326
2633842
20A9776D-AE5F-41BC-A35B-0C5E42EDFE48
Kolbasinella
Khramov, 2014a
Type
species.
Kolbasinella elongata
Khramov, 2014a
: 304
(by original designation).
Diagnosis.
Hind wing narrow; RP1 originating at approximately 1/5 wing length; RP with 12 branches MA pectinately branched near wing margin (7 closely spaced branches); numerous regularly arranged crossveins throughout wing, outer gradate series poorly formed near wing margin; MP with only a few pectinate branches; CuA strongly pectinate (
ca
. 19 very short branches); FW unknown.
Comments
.
Kolbasinella
was placed in
Gumillinae
by
Khramov (2014a)
due to the crossvein density and lack of trichosors in the wing margin. While these are indeed traits exhibited (in part) by
Gumillinae
, they are not diagnostic of the subfamily; some
Gumillinae
do have trichosors in the distal part of the wing (e.g.,
Gumilla
). Unfortunately, since
Kolbasinella
is only known from a hind wing, the justification presented by
Khramov (2014a)
for placement in
Gumillinae
is equivocal as important diagnostic antennal and forewing features are absent. The hind wing of
Kolbasinella
does not show the vein CuP but based on the shape of the other wing veins and the wing margin, it suggests that CuP is at least not elongate and pectinately branched, therefore placing it in one of three subfamilies,
Spilosmylinae
,
Protosmylinae
or
Gumillinae
. However, the genus is tentatively retained in
Gumillinae
here as its hind wing does not possess the basal sinuate crossvein
1r-m
, a state diagnostic for the subfamily (
1r-m
is present in
Spilosmylinae
and
Protosmylinae
). The lack of end-twigging in the posterior part of the wing is typical of
Gumillinae
but could also be due to the elongate shape of the wing and the proximity of the longitudinal veins (i.e., M and Cu) to the wing margin.