Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary pallgergely2@gmail.com Author Hunyadi, Andras Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary Author Vermeulen, Jaap J. JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands Author Grego, Jozef Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia Author Sutcharit, Chirasak Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Reischuetz, Alexander Puechhaimgasse 52, A- 3580 Horn, Austria Author Dumrongrojwattana, Pongrat Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand Author Botta-Dukat, Zoltan Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary Author Oerstan, Aydin 12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA Author Fekete, Judit University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary Author Jochum, Adrienne https://orcid.org/0000-0002-6624-6412 Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany text ZooKeys 2023 2023-02-13 1147 1 177 http://dx.doi.org/10.3897/zookeys.1147.93824 journal article http://dx.doi.org/10.3897/zookeys.1147.93824 1313-2970-1147-1 9BB9881B0076473D8E53155D37CA1F50 FF2B6B317B505F9EA0E1000BDCD16CE7 Angustopila steffeki Pall-Gergely & Grego sp. nov. Fig. 81 Type material. Holotype : Laos • 1 empty shell (H: 0.64 mm, D: 0.74 mm); Bolikhamsay Province, large spring lake at S foot of limestone massif 2 km NNE of Na Pavan village, ca. 5 km from Phontan crossing at road 8 (locality code: JG16); 18°13.30'N , 104°44.81'E ; 18 Feb. 2017; J. Grego leg.; NHMUK 20170296. Diagnosis. A small, depressed-globular Angustopila species with the aperture oblique to shell axis, a strongly protruding aperture, and relatively weak parietal and rather strong upper palatal tooth. Description. Shell small, wider than high; off-white, depressed-globular with domed spire; body whorl widest in standard apertural view; protoconch consists of 1.25 whorls, microstructure finely pitted and granular with a powdery superficial texture and with weak spiral striae (although holotype corroded; ca. 19 on body whorl from standard apertural view); teleoconch ornamented by some fine irregular, radial growth lines and stronger, equidistantly-arranged spiral striae; whorls 4, shouldered; aperture not oblique to shell axis from lateral view (although the peristome might be broken); umbilicus very wide; aperture pear-shaped with strongly distinct, relatively narrow sinulus and wider main section; peristome slightly expanded, not reflected; parietal callus strongly protruding, detached from penultimate whorl; upper parietal tooth relatively weak, low, but quite long, reaching peristome, perpendicular to parietal side; palatal tooth relatively strong, situated directly opposite of parietal tooth just at peristome edge. Figure 81. Angustopila steffeki Pall-Gergely & Grego, sp. nov. (holotype, NHMUK 20170296). Apertural ( A ), ventral ( B ), lateral ( C ) and apical ( D ) sides of the shell; sculpture of the protoconch ( E ); aperture ( F ); ventral ( G ) and frontal ( H ) surface of the body whorl. Measurements (in mm). H = 0.64, D = 0.74, H/D*100 = 86.5, RUD = 36.3 (holotype). Differential diagnosis. In comparison to other Angustopila species possessing two apertural denticles, Angustopila steffeki sp. nov. shows a more depressed shell with a dome-like appearance. Angustopila pallgergelyi has a more conical shell, a lower spire, and denser spiral striation. Angustopila somsaki sp. nov. has a less globular shell, less cylindrical (more funnel-shaped) umbilicus, and a less pointed palatal tooth. Etymology. This species is dedicated to and named after Jozef Steffek (1952-2013), Slovak malacologist. Distribution. This species is known from the type locality only in Bolikhamsay Province of Laos (Fig. 48 ).