Late Miocene Conidae (Mollusca: Gastropoda) of Crete (Greece). Part 1: genera Conilithes Swainson, 1840 and Conus (Kalloconus) da Motta, 1991 Author Psarras, Christos National and Kapodistrian University of Athens, Faculty of Geology & Geoenvironment, Department of Hist. Geology-Paleontology, Athens (Greece) cpsarras@geol.uoa.gr Author Koskeridou, Efterpi National and Kapodistrian University of Athens, Faculty of Geology & Geoenvironment, Department of Hist. Geology-Paleontology, Athens (Greece) ekosker@geol.uoa.gr Author Merle, Didier CR 2 P (CNRS, MNHN, UPMC, Sorbonne Université), Département Origines et Évolution, Muséum national d’Histoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) didier.merle@mnhn.fr text Geodiversitas 2021 2021-12-02 43 24 1309 1339 journal article 3058 10.5252/geodiversitas2021v43a24 d9b664cc-3197-44d5-92ca-08f8ce0fcc4f 1638-9395 5764710 urn:lsid:zoobank.org:pub:D36D1E14-73BE-4176-8024-F3673A65B8C1 Conus ( Kalloconus ) letkesensis ( Harzhauser & Landau, 2016 ) ( Figs 22 , 23 ; Table 10 ) Kalloconus letkesensis Harzhauser & Landau, 2016: 63 , figs 3M, 10E 1 -E 3 , 10F 1 -F 4 , 10G 1 -G 4 , 10H 1 -H 3 . Monteiroconus tietzei – Kovács & Vicián 2013 (partim): 79, figs 92, 94. TYPE MATERIAL. — Holotype NHMW 2016/0006/0001 . Paratype NHMW 2016/0006/0002 ). TYPE LOCALITY. — Letkés ( Hungary ) – Middle Miocene (Langhian) STRATIGRAPHIC RANGE. — Langhian of Paratethys (Pannonian Basin, Letkés, Hungary), Tortonian of Greece (Messara Basin, Crete). MATERIAL EXAMINED. — Greece . Filippi : 1 specimen AMPG(IV) 2682 . DESCRIPTION OF COLOUR PATTERN The colour pattern consists of multiple, closely placed spirals of long and short, very thin, fluorescent dashes. Dashes are not of constant length, with some resembling dots, whereas others resemble continuous spiral lines. Short dashes are usually between long dashes, while there are areas of the shell with multiple long dashes, there are areas that do not have any fluorescent colour ( Fig. 23 ). The colour pattern on the FIG. 21. — PCA graph with four species of Conus ( Kalloconus ) with similar colour pattern variations. Dots and area colors: ● , Conus ( Kalloconus ) hungaricus Hoernes & Auinger, 1879 ; ● , Conus ( Kalloconus ) tietzei Hoernes & Auinger, 1879 ; ● , Conus ( Kalloconus ) gulemani Erünal-ErentÖz, 1958 ; ● , Conus ( Kalloconus ) helladicus n. sp. ;  , best preserved specimens from both species described herein. Abbreviations: LW , length width ratio; RD , relative diameter; PMD , position of maximum diameter; RSH , relative height of spire. TABLE 10. — Shell measurements and ratios Conus ( Kalloconus ) letkesensis ( Harzhauser & Landau, 2016 ) from Filippi, Crete (Greece), AMPG(IV) 2682.

SL	MD	AH	HMD	AL	SA	LWA	LW	RD	PMD	RSH	SSFD	SSFd	PV
52.45 mm	35.45 mm	49.25 mm	42 mm	48.9 mm	150°	50°	1.48	0.72	0.85	0.06	2	9	0.29

TABLE 11. — Shell measurements and ratios of Conus ( Kalloconus ) asterousiaensis n. sp. from the Tortonian of Crete (Greece). 17 specimens measured; the largest specimen AMPG(IV) 2689 comes from Tefeli.

SL	MD	AH	HMD	AL	SA	LWA	LW	RD	PMD	RSH	SSFD SSFd		PV
Largest specimen	50.05 mm	32.2 mm	43.15 mm	37.5 mm	41.85 mm	110°	34°	1.55	0.75	0.87	0.14	2.73	4.57	0.23
Mean	28.2 mm	17.2 mm	24.3 mm	20.9 mm	24.4 mm	128.1°	37.5°	1.69	0.7	0.86	0.14	3.11	7.8	0.67
Standard deviation	10.52	7.47	9.36	8.36	9.48	16.03	3.81	0.18	0.07	0.03	0.04

spire whorls is partially destroyed on this specimen, but most likely consists of fluorescent flammulae ( Fig. 22 ). REMARKS This species has a low spire and a broad, conical last whorl, with smooth shoulder and a flat-sided last whorl ( Table 10 ). The colour pattern consists of delicate spiral rows of dashes. These characters are typical of Conus ( Kalloconus ) letkesensis ( Harzhauser & Landau, 2016 ) and therefore we attribute our specimen to this species. The Greek specimen is larger than the Paratethyan specimens (largest Paratethyan: 40.9 mm , versus Greek specimen: 52.45 mm ) and it displays a slightly different subsutural flexure (moderately curved in Harzhauser & Landau 2016 ). Harzhauser & Landau (2016) described three darker bands on the last whorl of the Paratethyan Conus ( Kalloconus ) letkesensis ( Harzhauser & Landau 2016 : fig. 10F-H). These bands are not visible on the Cretan specimen. The presence of this species in the Tortonian Eastern Protomediterranean, is an addition to the important cohort of taxa common to both the Middle Miocene of the Paratethys and eastern Proto-Mediterranean. The typical Paratethyan Langhian assemblages ( Harzhauser & Landau 2016 ) were found in the Proto-Mediterranean in the Serravallian of Turkey ( Landau et al. 2013 ). This species is an example of the persistence of some taxa into the late Miocene of Eastern Proto-Mediterranean.