Cabardites, a new genus for the “ Adites ” maculata (Poujade, 1886) species-group with descriptions of five new species from northern Indochina and eastern China (Lepidoptera: Erebidae: Arctiinae: Lithosiini) Author Volynkin, Anton V. 0000-0001-9447-4925 Altai State University, Lenina Av. 61, RF- 656049, Barnaul, Russia & National Research Tomsk State University, Lenina Av., 36, RF- 634050, Tomsk, Russia & monstruncusarctia @ gmail. com; https: // orcid. org / 0000 - 0001 - 9447 - 4925 monstruncusarctia@gmail.com Author Černý, Karel 0000-0001-7207-2255 Tiergartenstrasse 27, A- 6020 Innsbruck, Austria & natura. cerny @ aon. at; https: // orcid. org / 0000 - 0001 - 7207 - 2255 natura.cerny@aon.at text Zootaxa 2021 2021-01-25 4915 4 529 546 journal article 8513 10.11646/zootaxa.4915.4.4 1d51f348-b3b2-46ff-a370-636249acbdc3 1175-5326 4461583 C8CDB27B-1B5D-428A-9D13-0D95083C702C Cabardites limbata ( Wileman, 1911 ) , stat. rev. & comb. n. ( Figs 1–4 , 31, 32 , 47 ) Asura limbata Wileman, 1911 , The Entomologist 64 : 110 ( Type locality: [ Taiwan , Tainan , Guanziling ] “ Kanshirei ( 1000 ft. )”. = Parasiccia maculata var. formosibia Strand, 1917 , Archiv für Naturgeschichte 1916 (A3): 121 ( Type locality: [ Taiwan ] “Suisharyo … und … Taihorinsho”). = Parasiccia maculata f. shirakii Matsumura, 1930 , Insecta matsumurana 5 : 39 ( Type locality: [ Taiwan , Nantou County , Puli ] “ Formosa ; … at Horisha (?)”) . Type material examined . Lectotype of Asura limbata (designated herein) ( Fig. 1 ): male, “♁ Kanshirei , Formosa . 1,000 ft. 27.IV.1907 . A.E. Wileman ”/“ Asura limbata spec. nov. Type ♁”/“A 695 F”/“Wileman Coll. B.M. 1929- 261”/red ring “Type” label/QR-code label with a unique number “ NHMUK010598494 ” (Coll. NHMUK ) . Lectotype of Parasiccia maculata var. formosibia (designated herein) ( Figs 2 , 32 ): male, “ Kosempo ( Formosa ) H. Sauter X.1911 ”/“ Parasiccia maculata v. formosibia ♁ m. Strand det.”/red label “ Paratypus ♁ Paras. maculata v. formosibia Strd. ”/brown label “COTYPUS”/“ Milt. limbata Wilem. Wileman det.”/“ Milt. limbata Wilem. det. F. Daniel ”/“ Samml. F. Daniel ”/“Slide ZSM Arct. 2019-218♁ A. Volynkin ” (Coll. MWM / ZSM ) . Additional material examined . TAIWAN : 1 male , Prov. Hualien , Taroko N.P. , Hsipan , 270m , 5.IV.1997 , leg. Csorba & Ronkay , slide MWM 36212 Volynkin ( Coll. MWM / ZSM ) ; 1 male , Prov. Hua-Lien , 3 km W Jua-Juan , 150m , 02.VII.1996 , leg. Csorba & L. Németh ( Coll. MWM / ZSM ) ; 49 males , 1 female , Prov. Taitung , Chihpen Hot Springs , 400m , 6.IV., 9.IV.1997 , leg. Csorba & Ronkay ( Coll. MWM / ZSM ) ; 1 male , Prov. Taitung , 7 km E of Taimali , 710m , 4.VII.1997 , 22°35’N 120°53’E , leg. Cs ̂vari & Mikus ( Coll. MWM / ZSM ) ; 4 males , Prov. Taitung , 4 km N of Tupan , 390m , 14.VII.1997 , 22°28’N 120°52’E , leg. Tibor Csővári & László Mikus , slide MWM 36213 Volynkin ( Coll. MWM / ZSM ) ; 1 male , Lushan , Nantou , Formosa , 29.VIII.1973 , Y. Shibata / Inoue Coll. B.M. 1992-71/ NHMUK010605899 , slide NHMUK010313443 Volynkin ( Coll. NHMUK ) ; 1 female , Lushan Spa , 1200m , Nantou Hsien , Taiwan , 7–9.XI.1983 , Coll. K. Yazaki / Inoue Coll. B.M. 1992-71/ NHMUK010605901 , slide NHMUK010313444 Volynkin ( Coll. NHMUK ) ; 3 males , 3 females , Prov. Nantou , 17 km E of Kuohsing , 24°05’N 121°00’E , 560m , Ursun Forest Camping, 16. VI .1997 , leg. C. Szaboky & I. Soos ( Coll. MWM / ZSM ) ; 5 males , 1 female , Prov. Nantou , Huisun Forest Area , 500m , 15 km N Puli , 12–13.IV.1997 , leg. Csorba & Ronkay ( Coll. MWM / ZSM ) ; 1 male , Prov. Taipei , Hsiapen , 650m , 11.VII.1997 , leg. B. Herczig & Y.Y. Lien ( Coll. MWM / ZSM ) ; 1 female , Prov. Taipei , 10 km NE Pinglin , Pihou , 550m , 01. VI .1997 , leg. Gy. M. László & G. László ( Coll. MWM / ZSM ) ; 1 female , Prov. Taipei , 10 km SE of Pinglin , Pihou , 450m , 16.IV.1997 , leg. Gy. Fábián & S. T . Kovács , slide MWM 37049 Volynkin ( Coll. MWM / ZSM ) ; 1 female , Prov. Ilan , 1200m , Ming Chyr Forest Recreation Area , 8–9.VII.1997 , leg. B. Herczig & L. Ronkay ( Coll. MWM / ZSM ) ; 3 males , the same data, but leg. B. Herczig & S. Kovács ( Coll. MWM / ZSM ) ; 2 males , Prov. Kaohsiung , 700m , Liu-Kuei , San-Ping Forest Res. Stat., 22°58’16’’N 120°41’15’’E , 14–15.IV.1997 , leg. L. Peregovits & A. Kun ( Coll. MWM / ZSM ) ; 1 female , Prov. Kaohsiung , Tengchih Forest Recreation Area , 1200m , 19. V .1997 , leg. Gy. M. László & G. László ( Coll. MWM / ZSM ) ; 2 males , Prov. Ping-Tung , 5 km W of Chipen , 470m , 27–28.IV.1997 , leg. Gy. Fábián & S. Kovács ( Coll. MWM / ZSM ) ; 1 male , Pintung Hsien , Kenting Park , 100m , 26–27.III.1984 , A. Kawabe leg./ Inoue Coll. B.M. 1992-71/ NHMUK010605900 ( Coll. NHMUK ) ; 1 male , Prov. Taichung , Hui Sun Exp. Forest , Guandashi LTER site, 950m , 24°04’49’’N 121°02’08’’E , 12–13.IV.1997 , leg. Peregovits & Kun ( Coll. MWM / ZSM ) ; 1 male , Prov. Taoyuan , Ming Chyr Forest Recreation Area , 1160m , 17–18.IV.1997 , leg. Gy. Fábián & S. T . Kovács ( Coll. MWM / ZSM ) . Remarks . (1) The senior author dissected the syntype specimen of Parasiccia maculata var. formosibia Strand, 1917 deposited in MWM/ZSM and found no genital differences from other forms of C. limbata . To avoid any probable confusion of this taxon with the externally similar and also variable C. pica , here we designate the syntype dissected as lectotype . (2) Unfortunately, we did not find pictures of the holotype female of Parasiccia maculata f. shirakii Matsumura, 1930 among the photo materials of the Matsumura collection (EIHU, Sapporo) kindly provided by Dr. V.V. Zolotuhin Dr. V.V. Dubatolov. In his paper, Hiroshi Inoue (1988) also did not mention the type and established the synonymy without any comments. Taking these facts into account, we can only assume that the holotype is lost. However, the original description of this taxon given by Matsumura (1930) clearly corresponds to the external appearance of C. limbata therefore we consider shirakii as a junior synonym of C. limbata . (3) Wileman (1911) described Asura limbata from two specimens of both sexes, which are syntypes . To avoid any confusion in the future here we designate the male specimen labeled as “Type” as lectotype . FIGURES 1–10. Cabardites spp.: adults. Depositories of the specimens: 1, 5, 6 in NHMUK (©); 2–4 and 7–10 in MWM/ ZSM. FIGURES 11–20. Cabardites spp.: adults. Depositories of the specimens: 11, 12, 15, 17–20 in MWM/ZSM (18 and 19 – ex coll. CKC); 13 in MNHN (©); 14, 16 in NHMUK (©). FIGURES 21–30. Nudariina spp.: type species of the genera (21–24, 27–30) and non-type species (25, 26). Depositories of the specimens: 21, 22 in MWM/ZSM; 23, 24, 27, 28 in NHMUK (©); 25, 26 in CKC; 29, 30 in CAV. FIGURES 31–34. Cabardites spp.: male genitalia. The specimens are deposited in MWM/ZSM. Diagnosis . Forewing length is 9–10.5 mm in males and 11–12 mm in females. The species is similar externally to C. maculata , but can be easily distinguished by its shorter lamellae of male antennae, smaller size, the smoothly curved postmedial line lacking a deep concavity at costa and anal margin and two short triangular protrusions opposite the cell, and the paler hindwing with a weaker brownish suffusion. The male genitalia differ clearly from those of other congeners by the aedeagus vesica bearing a series of very long, robust spines. The differences between C. limbata and its closest relative C. pica are listed in the diagnosis of the latter species. Distribution . Endemic to Taiwan Island .