<document id="103C56D6C1B4C898AC85361BD5767103" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong &amp; Prokop, Jakub" docDate="2012" docId="B970A055FFCFFFEAFED1252DFC79FBAC" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Helophorus (Mesosperchus) inceptivus Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="104" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="104" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3A4B30FAE2DBFCC3BDBD229071C17591">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
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<mods:namePart id="C48FC0EFEB29965664D905E654F63E10">Fikáček, Martin</mods:namePart>
<mods:affiliation id="9DF22772A7E0B988727AD4B315E58C81">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="9C1E75B62A43C182313A5F5B6D491021">Prokin, Alexander</mods:namePart>
<mods:affiliation id="C5FF6EFE3BE4AAE2300910AB81F0E837">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru &amp; Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
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<mods:namePart id="4ABAC2ECE918818A2E6480BF025664A8">Angus, Robert B.</mods:namePart>
<mods:affiliation id="EFAEDD54A1BE583570A994161DEE5C2F">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
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<mods:namePart id="2EFD872DDE3365F147FBBDA39A70447C">Pono, Alexander</mods:namePart>
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<mods:namePart id="8A39854A0002BD8E55A1145D2AB7141F">Marenko</mods:namePart>
<mods:affiliation id="C1CB184642D7EE9B7BDD69AF46AC7F50">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
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<mods:namePart id="13568825D43F8B9DB6BD1714F51C8593">Yue, Yanli</mods:namePart>
<mods:affiliation id="0D6675D66A960725A8AA1A058C8798CE">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
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<mods:namePart id="09CFECD53962CAC15DC2B11ED028B7D7">Ren, Dong</mods:namePart>
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<mods:namePart id="94D58A26679C8453BEDDAF7FD8B6F376">Prokop, Jakub</mods:namePart>
<mods:affiliation id="2F3DF59ED4FFE37D87117E1D79BD101A">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:title id="81CF946B681B44200DACF4FFEA708E6C">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DD4A528F838830DBB6A0A71CBA7E64F9">2012</mods:date>
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<heading id="6A2EA62FFFCFFFEAFED1252DFCD1FD6F" bold="true" box="[330,872,635,660]" centered="true" fontSize="8" level="2" pageId="15" pageNumber="104" reason="3">
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† 
<taxonomicName id="F6D96AC0FFCFFFEAFEC5252DFCB5FD68" ID-CoL="3KHKX" authority="Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop, 2012" authorityName="Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop" authorityYear="2012" box="[350,780,635,660]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="inceptivus" status="sp. nov." subGenus="Mesosperchus">
<emphasis id="03ADCD51FFCFFFEAFEC5252DFE59FD6F" bold="true" box="[350,480,635,660]" italics="true" pageId="15" pageNumber="104">Helophorus</emphasis>
(
<emphasis id="03ADCD51FFCFFFEAFE6B252DFD34FD6F" bold="true" box="[496,653,635,660]" italics="true" pageId="15" pageNumber="104">Mesosperchus</emphasis>
) 
<emphasis id="03ADCD51FFCFFFEAFD06252DFCB5FD68" bold="true" box="[669,780,635,659]" italics="true" pageId="15" pageNumber="104">inceptivus</emphasis>
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<taxonomicNameLabel id="189E702AFFCFFFEAFC89252AFCD1FD6F" box="[786,872,636,660]" pageId="15" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="79C342C8FFCFFFEAFE7D25CCFD75FD56" box="[486,716,666,685]" pageId="15" pageNumber="104" type="description">
<paragraph id="31661143FFCFFFEAFE7D25CCFD75FD56" blockId="15.[486,716,666,685]" box="[486,716,666,685]" pageId="15" pageNumber="104">
(
<figureCitation id="A9E20DC6FFCFFFEAFE7725CCFD8AFD56" box="[492,563,666,685]" captionStart="Figs" captionStartId="16.[149,188,896,915]" captionTargetBox="[232,1072,226,879]" captionTargetId="figure-242@16.[206,1072,216,884]" captionTargetPageId="16" captionText="Figs. 27–28. Representatives of Helophorus, subgenus Mesosperchus Ponomarenko, 1977. 27 – H. (M.) inceptivus sp. nov. (PIN 4270/1008, holotype); 28 – H. (M.) yixianus sp.nov. (CNU 2010004, holotype). Scale bars: 1 mm. Abbreviations: ant – antenna." figureDoi="http://doi.org/10.5281/zenodo.5330630" httpUri="https://zenodo.org/record/5330630/files/figure.png" pageId="15" pageNumber="104">Figs. 27</figureCitation>
, 
<figureCitation id="A9E20DC6FFCFFFEAFDA725CCFDCAFD56" box="[572,627,666,685]" captionStart="Figs" captionStartId="25.[113,152,1499,1518]" captionTargetBox="[123,1078,217,1475]" captionTargetId="figure-15@25.[123,1078,217,1476]" captionTargetPageId="25" captionText="Figs. 73–82. Representatives of Helophorus, subgenus Mesosperchus Ponomarenko, 1977. 73–79 – Helophorus tarsalis (Ponomarenko, 1977) (73 – PIN 3063/116, paratype; 74 – PIN 3063/2192, piece, general view; 75 – PIN 3015/1769; 76 – PIN 3063/2192, counterpiece, detail of male genitalia; 77– PIN 3063/2194. piece under alcohol; 78 – same specimen, counterpiece in dry condition; 79 – PIN 3015/367, holotype). 80 – Helophorus gracilis (Prokin, Ren &amp; Fikáček, 2010), CNU 2010018, holotype. 81–82 – Helophorus inceptivus, PIN 4270/1008 (81 –detail of head and pronotum; 82 – general view). Scale bars: 1 mm (Figs. 73–80, 82); 0.5 mm (Fig. 81)." figureDoi="http://doi.org/10.5281/zenodo.5330649" httpUri="https://zenodo.org/record/5330649/files/figure.png" pageId="15" pageNumber="104">81–82</figureCitation>
, 
<figureCitation id="A9E20DC6FFCFFFEAFDE725CCFD7FFD56" box="[636,710,666,685]" captionStart="Figs" captionStartId="29.[113,153,1497,1516]" captionTargetBox="[133,1068,219,1485]" captionTargetId="figure-15@29.[133,1069,219,1486]" captionTargetPageId="29" captionText="Figs. 104–112. Mesozoic representatives of the helophorid lineage, scanning electron micrographs. 104–106 – Laetopsia baissensis (Ponomarenko, 1986), PIN 3064/7018 (104 – general view; 105 – detail of head and pronotum; 106 – detail of meso- and metathorax). 107, 111 – Helophorus (Mesosperchus) inceptivus sp. nov. (107 – general view; 111 – detail of pronotum). 108–112 – Helophorus (Mesohelophorus) paleosibiricus nom. nov. (108 – PIN 3064/6981, general view; 109 – PIN 3063/863, holotype, 110 – PIN3064/6981, detail of prothorax, ventral view; 112 – same, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5330657" httpUri="https://zenodo.org/record/5330657/files/figure.png" pageId="15" pageNumber="104">107, 111</figureCitation>
)
</paragraph>
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<subSubSection id="79C342C8FFCFFFEAFFEA2592FD72FCDC" pageId="15" pageNumber="104" type="materials_examined">
<paragraph id="31661143FFCFFFEAFFEA2592FD92FCF7" blockId="15.[113,1089,707,807]" pageId="15" pageNumber="104">
<emphasis id="03ADCD51FFCFFFEAFFEA2592FE85FD2C" bold="true" box="[113,316,707,727]" pageId="15" pageNumber="104">
<typeStatus id="EE62AFE1FFCFFFEAFFEA2592FF27FD2C" box="[113,158,708,727]" pageId="15" pageNumber="104">Type</typeStatus>
locality and age.
</emphasis>
<collectingCountry id="49CE51D3FFCFFFEAFED82592FE21FD2C" box="[323,408,708,727]" name="Mongolia" pageId="15" pageNumber="104">Mongolia</collectingCountry>
, 
<collectingRegion id="F31DDFA1FFCFFFEAFE392592FDEAFD2C" box="[418,595,708,727]" country="Mongolia" name="Govi-Altay" pageId="15" pageNumber="104">Gobi-Altai Province</collectingRegion>
, SE Ajbogd-uul Range, S of Mount Shar-Teg, fossil site Shar-Teg, exposure 442/2; Shar-Teg sequence, Late Jurassic, Tithonian, ca. 150–145 mya (
<bibRefCitation id="55486CB2FFCFFFEAFCE72588FB84FD0A" author="GUBIN Y. M. &amp; SINITZA S. M." box="[892,1085,734,753]" pageId="15" pageNumber="104" pagination="311 - 318" refId="ref14967" refString="GUBIN Y. M. &amp; SINITZA S. M. 1996: Shar Teg: a unique Mesozoic locality of Asia. In: MORALES M. (ed.): The Continental Jurassic. Museum of Northern Arizona Bulletin 60: 311 - 318." type="journal article" year="1996">
<collectionCode id="57C88986FFCFFFEAFCE72588FC09FD0B" box="[892,944,734,753]" pageId="15" pageNumber="104">GUBIN</collectionCode>
&amp; SINITZA 1996
</bibRefCitation>
; 
<bibRefCitation id="55486CB2FFCFFFEAFFEA25AFFED5FCF7" author="RASNITSYN A. P. &amp; ZHERIKHIN V. V." box="[113,364,761,780]" pageId="15" pageNumber="104" pagination="437 - 444" refId="ref15948" refString="RASNITSYN A. P. &amp; ZHERIKHIN V. V. 2002: Alphabetic list of selected insect fossil sites - Impression fossils. Pp. 437 - 444. In: RASNITSYN A. P. &amp; QUICKE D. L. J. (eds.): History of Insects. Kluwer Academic Publishers, Norwell, 517 pp." type="book chapter" year="2002">RASNITSYN &amp; ZHERIKHIN 2002</bibRefCitation>
; 
<bibRefCitation id="55486CB2FFCFFFEAFEEF25AFFD99FCF7" author="YAN E. V. &amp; ZHANG H. C." box="[372,544,761,780]" pageId="15" pageNumber="104" pagination="451 - 456" refId="ref16203" refString="YAN E. V. &amp; ZHANG H. C. 2010: New beetle species of the formal genus Artematopodites (Coleoptera: Polyphaga), with remarks on the taxonomic position of the genera Ovivagina and Sinonitidulina. Paleontological Journal 44: 451 - 456." type="journal article" year="2010">
YAN &amp; 
<collectionCode id="57C88986FFCFFFEAFE2E25AFFE54FCF0" box="[437,493,761,780]" pageId="15" pageNumber="104">ZHANG</collectionCode>
2010
</bibRefCitation>
).
</paragraph>
<paragraph id="31661143FFCFFFEAFFEA2445FD72FCDC" blockId="15.[113,1089,707,807]" box="[113,715,787,807]" pageId="15" pageNumber="104">
<emphasis id="03ADCD51FFCFFFEAFFEA2445FEA7FCDC" bold="true" box="[113,286,787,807]" pageId="15" pageNumber="104">Material examined</emphasis>
(
<specimenCount id="27DFDACAFFCFFFEAFEB02442FED3FCDC" box="[299,362,788,807]" pageId="15" pageNumber="104" type="generic">1 spec.</specimenCount>
). 
<typeStatus id="EE62AFE1FFCFFFEAFEEC2442FE72FCDE" box="[375,459,788,807]" pageId="15" pageNumber="104" type="holotype">HOLOTYPE</typeStatus>
: PIN 4270/1008 (piece only).
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFCFFFEAFFEA2469FC79FBAC" pageId="15" pageNumber="104" type="description">
<paragraph id="31661143FFCFFFEAFFEA2469FC79FBAC" blockId="15.[113,1089,830,1111]" pageId="15" pageNumber="104">
<emphasis id="03ADCD51FFCFFFEAFFEA2469FEFEFCAC" bold="true" box="[113,327,830,855]" pageId="15" pageNumber="104">
Description. 
<emphasis id="03ADCD51FFCFFFEAFE9A2468FE87FCAC" bold="true" box="[257,318,830,855]" italics="true" pageId="15" pageNumber="104">Adult</emphasis>
:
</emphasis>
Body rather wide, body length 
<quantity id="F621BCA6FFCFFFEAFD022469FD57FCAC" box="[665,750,831,855]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="15" pageNumber="104" unit="mm" value="3.9">3.9 mm</quantity>
. Frons partly bearing weak granulation, gula wide, antenna with three-segmented antennal club. Pronotum transverse, with simply arcuate lateral margins and slightly angulate posterior margin; surface bearing five longitudinal grooves, all pronotal intervals with well-developed (through mostly only weakly preserved) granulation; anterolateral corners of pronotum probably not projecting. Prothoracic antennal grooves probably present. Elytron with 10 series of punctures, all series finely impressed; elytral intervals flat. Mesocoxal cavities very narrowly separated from each other, metaventrite slightly longer than mesoventrite. Mesotibia with several longitudinal series of spines, bearing few larger spines on distal margin; abdomen with five ventrites.
</paragraph>
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