A revision of the genera Pseudosmittia Edwards, 1932, Allocladius Kieffer, 1913, and Hydrosmittia gen. n. (Diptera: Chironomidae, Orthocladiinae) 2849
Author
Ferrington Jr, Leonard C.
Author
Saether, Ole A.
text
Zootaxa
2011
2011-04-29
2849
1
1
314
https://biotaxa.org/Zootaxa/article/view/zootaxa.2849.1.1
journal article
10.11646/zootaxa.2849.1.1
11755334
5288200
Pseudosmittia nishiharaensis
Sasa
et
Hasegawa
(
Fig. 72 A–E
)
Pseudosmittia nishiharaensis
Sasa
et
Hasegawa, 1988: 247
;
Saether 2006: 39
;
Makarchenko & Makarchenko 2008: 221
.
Pseudosmittia linguata
Caspers
et
Reiss, 1989: 128
;
Saether & Ferrington, 2003: 5
.
Pseudosmittia yakymenea
Sasa et Suzuki, 2000a: 92
;
Yamamoto 2004: 90
.
Pseudosmittia yakyneoa
Sasa et Suzuki, 2000a: 93
;
Yamamoto 2004: 90
.
Material examined
:
JAPAN
: Ryukyus,
Okinawa Pref.
, Miyako Island, Nishihara,
holotype
male,
3.ii.1982
, M. Sasa & H. Hasegawa (NMST A 65:22); Ryukyus,
Okinawa Pref.
, Miyako, Ikema, and
Okinawa
(Kochinda, Yara Bridge, Kadena-cho) Islands,
paratypes
11 male
, 2 doubtful females, including 2 misidentified males,
11.xii.1981
,
28.i.1982
,
3.ii.1982
, M. Sasa & H. Hasegawa [NMST A 65:13 (misidentified
P. topei
), 19–21, 24, 83–84];
Kagoshima Pref.
, Osumi Islands, Yakushima Island, Miyanoura, Issogawa,
holotype
male of
P. yakymenea
,
holotype
male of
P. yakyneoa
,
23.iii.1999
, H. Suzuki (NMST;
holotype
of
P. yakymenea
: No.
382: 27,
holotype
of
P. yakyneoa
: No.
382: 28).
CHINA
:
Ningxia
, Mt. Linpan,
3 males
,
4.vii.1984
, X. Wang.
THAILAND
: Chiangmai Zoo,
2 males
,
18–30.vi.1988
,
27.ii.–6.iii.1989
, H. Malicky (
ZSM
); Erzurum Prov., Azort Pass west of Oltu,
2200 m
a.s.l.,
holotype
male of
P
.
linguata
,
6.vii.1985
(
ZSM
).
ITALY
:
Lombardy
, Angera, Lago Maggiore,
paratype
male of
P
.
linguata
,
31.vii.–6.viii.1965
, F. Reiss (
ZSM
).
Diagnostic characters
: The presence of a strong apical antennal seta combined with a forked postcubitus, AR of 0.9–1.4, and the apically truncate and upcurved gonostylus separate this species from other members of the group.
Male imago
(n = 11–12, except when otherwise stated)
All measurements and ratios within the range of variation of
P
.
forcipata
with the following exceptions and additions: Wing (
Fig. 72 A
) length/length of profemur 2.86–3.29, 3.07. AR 0.88–1.43, 1.06. Clypeus with 4–6, 5 (7) setae. Palpomere lengths (in µm): 18–26, 21; 23–34, 28; 48–56, 53; 67–75, 71; 80–117, 102. Third palpomere (
Fig. 72 C
) with 2 (7) sensilla clavata at apex. VR 1.60–1.85, 1.69; VR
2
1.30–1.46, 1.37. Costal extension (
Fig. 79 F
) 0–25, 16 µm long. R
4+5
ending well proximal to apex of M
3+4
, Cu
1
slightly curved to straight. LR
2
0.50–0.55, 0.53; BV
2
3.78–4.15, SV
2
3.55–3.87,
BR
3
7.3–8.4. Anal point (
Fig. 72 F
;
Sasa & Hasegawa 1988
, fig. 6j;
Caspers & Reiss 1989
, fig. 17;
Sasa & Suzuki 2000a
, fig. 52i) 30–49, 41 µm long, with 6–13, 7 setae on sides and at base. Phallapodeme 51–60, 55 µm long. Virga (
Fig. 72 E
) 14–21, 16 µm long. Gonocoxite with inferior volsella reaching to 0.64–0.85, 0.72 gonocoxite length.
Female
Possibly known (see below).
Pupa and larva
Unknown.
Remarks
The female imago is partly described by
Sasa and Hasegawa (1988: 248)
. However, the
paratype
females are tentatively associated, damaged, possibly belong to different species, two or three dissected specimens are placed under the same cover slip, and the wings are dry mounted making it impossible to obtain an accurate count on the nonmarginal setae on the costa. A redescription thus is not warranted on the present material.
Both
Sasa and Hasegawa (1988)
,
Caspers and Reiss (1989)
and
Sasa and Suzuki (2000a)
overlooked the postcubital forking which is indistinct and difficult to distinguish in most specimens. The anal point in
Sasa and Suzuki (2000a)
is wrongly drawn.
Distribution
The species is known from
Germany
,
Italy
,
Turkey
, Russian Far East,
China
,
Japan
including Indo-Pacific parts, and
Thailand
.