Redescription of Endonura taurica (Stach, 1951) and E. quadriseta Cassagnau & Péja, 1979, and description of two new species of the genus Endonura Cassagnau, 1979 (Collembola: Neanuridae: Neanurinae) from the Crimea (Ukraine) Author Smolis, Adrian Author Skarżyński, Dariusz Author Pomorski, Romuald J. Author Kaprus’, Ighor J. text Zootaxa 2007 1442 19 35 journal article 10.5281/zenodo.176031 6a062adb-aa9f-42d9-8bb1-6de6a10e8806 1175-5326 176031 Endonura dentifera sp. nov. Figs 38–49 , Tab. 4 Material examined . Type material. Holotype , adult female on slide, leaf litter on mountain meadow, Nikits’kyi Pereval, ca. 1450 m a.s.l., neighborhood of Jalta, Crimea, Ukraine , 12.ix.1997 , leg. R. J. Pomorski, D. SkarŻyński, I. Kaprus’. Paratypes : 3 males , 2 females and juvenile on slides, the same data as holotype ( holotype and 2 paratypes ZIWU , 4 paratypes MNHL ). TABLE 3. Chaetotaxy of Endonura quadriseta . a) Cephalic chaetotaxy.

Tubercle	Number of chaetae	Types of chaetae	Names of chaetae
Cl Af	4 11	Ml Mc Ml Mc	F G B A,O,C, D,E,
Oc	3	Ml Mc me	Ocm Ocp Oca
Di	2	Ml Mc	Di1 Di2
De Dl	2 6	Ml Mc Ml Mc	De1 De2 Dl1,Dl5 Dl2–4,Dl6
(L+So)	9	Ml Mc me	L1,L4,So1 L2,L3 So3–6

Number of other cephalic chaetae: Vi, 6; Ve, 10–11; labrum, 4/2, 4; labium, 11, 0x; ant. I, 7; ant.II, 11; ant. III, 18+5s; ant. IV, 8S+i +or+12mou. b) Postcephalic chaetotaxy.

th. I	Di 1	Terga De 2	Dl L 1 –	Scx2 –	Cx 3	Legs Tr 6	Fe 13	T 19
th. II	3	2+s	3+s+ms 3	2	7	6	12	19
th. III	3	3+s	3+s 3	2	8	6 Sterna	11	18
abd. I	2	3+s	2 3	VT: 4

abd. II 2 abd. III 2	3+s 2 3+s 2	3 4	Ve: 5 Ve1 present Ve: 5–6	Fu: 4–7me 0 mi
abd. IV 2	2+s 4	6–8	Ve: 8	Vl:4
abd. V (3+3) abd. VI	6–7+s 7	Ag: 3 Ve: 11–15	Vl: 1 L’:1 An: 2 mi

Diagnosis . Habitus typical of the genus Endonura . Dorsal tubercles present, well developed. 2+2 eyes present. Buccal cone elongated. Labral chaetotaxy 4/2, 4. Mandible thin with 5 teeth. Head with 3 chaetae Oc, chaetae A, B, C, D, E (chaeta O absent). Tubercle (L+So) on head with 10 chaetae. Tubercles De on thoracic terga II and III with 3 and 4 chaetae respectively. Tubercles L on abd. III and IV with 4 and 8–9 chaetae respectively. Claw with inner tooth. Tibiotarsi I, II with long chaetae B5 and B4. Tibiotarsus III with only one long chaeta B4. Etymology . The species name is derived from the Latin words “dens”–tooth and “fero”–wear. The new species has a claw with distinct internal tooth. Description . Habitus typical of the genus. Body length (without antennae) 1.45–1.95 mm. Colour of the body greyish-blue of variable intensity. 2+2 large, dark-pigmented eyes. FIGURES 27–32 . Endonura quadriseta : 27, habitus and dorsal chaetotaxy (holotype); 28, chaetotaxy of tubercles (So+L) and Oc on head; 29, ventral sclerifications of labrum; 30, chaetotaxy of labium and group Vi; 31, microsensillum and ventral guard sensillum in antennal III-organ; 32, antennal segments III–IV of right antenna (dorsal view). FIGURES 33–37 . Endonura quadriseta : 33, tibiotarsus and claw III (dorsolateral view); 34, tibiotarsus and claw III (ventrolateral view); 35, chaetae of male ventral organ; 36, ventral chaetotaxy of abd. II–VI (adult male); 37, chaeta Di1 of abd. V. Types of dorsal ordinary chaetae: macrochaetae Ml and Mc thickened, narrowly sheathed, feebly serrated and not pointed ( Figs 38, 42–44 ); mesochaetae thin and pointed. Head. Buccal cone rather long and rounded at apex. Labrum chaetotaxy 4/2, 4, ventral sclerifications nonogival. Chaetotaxy of labium as in Fig. 40 and Tab. 4 . Maxilla styliform, mandible thin with 2 basal and 3 apical teeth ( Fig. 39 ). Chaetotaxy of antennae as in Tab. 4 . Apical vesicle distinct, trilobed. S-chaetae subequal, long and moderately thickened. Chaetotaxy of head as in Figs 38, 40–41 and Tab. 4 . Tubercle Af narrow ( Fig. 38 ), chaeta O absent. Tubercle (L+So) with 10 chaetae (chaeta So2 present, Fig. 41 ). Thorax, abdomen, legs. Chaetotaxy of th. and abd. as in Figs 42, 44–45 and Tab. 4 . Tubercles Di on th.I not differentiated. Tubercles L on abd. III and IV with 4 and 8–9 chaetae respectively ( Fig. 45 ). Cryptopygy absent. Chaetotaxy of legs as in Figs 46–49 and Tab. 4 . Claw with distinct inner tooth ( Figs 46–49 ). Tibiotarsi I and II with elongate and slightly clavate chaetae B5 and B4. Tibiotarsi III with elongate and slightly clavate chaeta B4. Discussion . The new species is the most similar to E . quadriseta Cassagnau et Péja, 1979 , but differs from it in the following characters: tubercle (L+So) on head with chaeta So2 (in quadriseta absent), chaeta O absent (in quadriseta present), claw with inner tooth (in quadriseta absent) and tibiotarsi with elongate and slightly clavate chaetae B4 and B5 (in quadriseta short and not clavate). TABLE 4 . Chaetotaxy of Endonura dentifera sp. nov. a) Cephalic chaetotaxy.

Tubercle	Number of chaetae	Types of chaetae	Names of chaetae
Cl Af	4 10	Ml Mc Ml Mc	F G B A,C,D,E
Oc	3	Ml Mc me	Ocm Ocp Oca
Di	2	Ml Mc	Di1 Di2
De Dl	2 6	Ml Mc Ml Mc	De1 De2 Dl1,Dl5 Dl2–4,Dl6
(L+So)	10	Ml Mc Mc or me me	L1,L4,So1 L2 L3,So2 So3–6

Number of other cephalic chaetae: Vi, 6; Ve, 10–11; labrum, 4/2, 4; labium, 11, 0x; ant. I, 8; ant.II, 11; ant. III, 18+5s; ant. IV, 8S+i +or+12mou. b) Postcephalic chaetotaxy.

Di	Terga De	Dl L	Scx2	Cx	Legs Tr	Fe	T
th. I	1	2	1 –	–	3	6	13	19
th. II th. III	3 3	2+s 3+s	3+s+ms 3 3+s 3	2 2	7 8	6 6	12 11	19 18

Sterna abd. I 2 3+ s 2 3 VT: 4 abd. II 2 3+ s 2 3 Ve: 5–6 Ve1 present