Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae)
Author
Páll-Gergely, Barna
31E196E9-5A51-4295-9A36-D5DA689502B7
Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Ottó Street 15, Budapest, H- 1022, Hungary.
pall-gergely.barna@agrar.mta.hu
text
European Journal of Taxonomy
2018
2018-08-16
455
1
114
journal article
22297
10.5852/ejt.2018.455
44a2fcf6-cc1b-4c07-9554-dd6dd279a76d
3817707
C445E95B-446A-4601-AAA3-C1CCBAB627F9
Subfamily
Sinicolinae
subfam. nov.
urn:lsid:zoobank.org:act:
7AA09CFF-D5FD-4DB1-98BE-9DB9E41F6B82
Type
genus
Sinicola
Gude, 1899
.
Included genera
Gudeodiscus
Páll-Gergely, 2013
(with the subgenus
Veludiscus
Páll-Gergely, 2015
),
Endothyrella
,
Halongella
Páll-Gergely, 2015
,
Sicradiscus
Páll-Gergely, 2013
,
Sinicola
.
The Eocene fossil genus
Plectopyloides
Yen, 1969
(
type
species:
P. cretaceous
Yen, 1969
; synonyms:
Henanspirus
Li, 1986
,
Menyinspirus
Li, 1986
,
Yenellus
Li, 1986
) is also classified in this subfamily, because it is most similar to
Gudeodiscus
(see
Páll-Gergely & Asami
2014
).
Diagnosis
Shell usually dextral (although most species of
Endothyrella
are sinistral), flat to conical-globular; umbilicus wide to narrow; protoconch regularly ribbed without any additional sculptural elements (very rarely smooth, see
Páll-Gergely
et al.
2015
b
,
2015
c); apertural fold present or absent; main plica usually absent (present only in
Gudeodiscus longiplica
Páll-Gergely & Asami, 2016
). Penis well-developed; penial caecum well-developed to absent; epiphallus present; diverticulum as long as bursa copulatrix (absent in some species, although possibly overlooked during dissection).
Distribution
Same as in the family
Plectopylidae
, except
Thailand
and the Malay Peninsula (
Fig. 3
).
Remarks
The subdivision of the family into two reciprocally monophyletic groups is supported by molecular phylogenetic analysis, which included nearly all genera. Discussion of the phylogenetic results is beyond the scope of the present revision, and will be discussed in a separate paper.