A review of the late Cenozoic genus Bohra (Diprotodontia: Macropodidae) and the evolution of tree-kangaroos Author Prideaux, Gavin J. 0000-0002-9958-0265 gavin.prideaux@flinders.edu.au Author Warburton, Natalie M. 0000-0002-9958-0265 gavin.prideaux@flinders.edu.au text Zootaxa 2023 2023-06-07 5299 1 1 95 http://dx.doi.org/10.11646/zootaxa.5299.1.1 journal article 53706 10.11646/zootaxa.5299.1.1 4aabf565-7d02-4801-a31c-62415ccee45f 1175-5326 8012666 9CA85AEC-7128-4118-A50D-FCD16502F5E0 Bohra sp. cf. B. bandharr cf. Dendrolagus sp. 1 : Flannery & Archer (1984) , pp. 358–359, 361–362, figure 1F–G, table 1. Rich (1991) , p. 1049. cf. Dendrolagus sp. 2 : Flannery & Archer (1984) , pp. 358–359, 362–363, figure 1F–G, table 1. Rich (1991) , p. 1049. Tedford et al. (1992) , p. 187. cf. Dendrolagus : Tedford et al. (1992) , p. 190, table 3; partim. Protemnodon ?cf. bandharr : Dawson et al. (1999) , pp. 281, 287, table 10. Dendrolagus sp. 2 : Dawson et al. (1999) , p. 281 . Dendrolagus sp. : Dawson (2004a) , pp. 267, 270–272, table 1; partim. Silvaroo sp. indet. 1: Dawson (2004b) , pp. 283, 286, figure 3. Silvaroo sp. 1 : Dawson (2004b) , p. 288 , table 2. Dendrolagus sp. indet.: Prideaux & Warburton (2008) , pp. 463–464, figure 1; partim. Silvaroo sp. : Louys & Price (2015) , pp. 565, 567, 569, figure 8N. Referred specimens. Chinchilla Rifle Range (site not specified), Queensland . QM F43281, partial right adult dentary (preserving m3–4). QM F58665, partial right adult dentary (preserving m3–4); QM F61040, right i1. Collected by Doris and Cecil Wilkinson. Probably Chinchilla. QM F4750, partial right adult dentary (preserving incomplete p3, m1–4). This specimen retains no associated locality information. Chinchilla Sand is suggested by preservation (chocolate-brown bone, blue-grey enamel and yellow- to orange- to chocolate-brown adhering matrix), which has been used to retrospectively, albeit tentatively, differentiate Chinchilla Sand specimens from other Darling Downs material (e.g., Bartholomai 1963 , 1975 ; Bartholomai & Woods 1976; Louys & Price 2015 ). Collection details are unknown, but it was very likely collected in the late 19 th century. The specimen was ascribed the number 8808 in De Vis’s catalogue. Bow, New South Wales . AM F59576, partial right adult maxilla (preserving M2–3); AM F64004, posterior half of left P3; AM F64007, right M4; AM F64075, partial left maxilla (preserving very worn M2–4). AM F64245, left M4. Fisherman’s Cliff, Warrananga Station, New South Wales . NMV P38267, left M2; MNV P38351, left M3 metaloph; NMV P38325, right p3. Collected by Edmund Gill and others in the late 1960s or James Warren and others in the early 1970s. FUPS Quarry , Bone Gulch , Moorna Station , New South Wales . NMV P257273 , partial left P3 ; NMV P257274 , left M3 ; NMV P257275 , right M1 metaloph. These specimens were picked from concentrate sieved from sediment collected by a team led by GJP . NMV P257273 was collected on 14 February 2016 , NMV P257274 on 16 August 2015 , and NMV P257275 on 15 February 2016 . Remarks. An assortment of specimens have been grouped under this limbo nomen ( Figure 16–18 ) because they share dentary or cheek-tooth traits with specimens of B. bandharr (e.g., comparatively rectangular upper molars, dentary with a long, rounded postalveolar shelf), but differ by having narrower, much smaller molars ( Table 2 ). NMV P38325, by contrast, is a p3 from the early Pleistocene Fisherman’s Cliff LF ( Figure 1 ) referred to B. sp. cf. B. bandharr by association with several loose molars, and the general similarity of its form ( Figure 18J–L ) to that of the only other unworn p3 known for a species of Bohra , that of the paratype of B. illuminata ( Figure 24D ). The barrier to defining a species from among this collective is the lack, in any one specimen , of sufficient diagnostic attributes that would permit its selection as a reliable holotype . Obtaining further tree-kangaroo fossils from the Chinchilla and Bow localities ( Figure 1 ) may offer the greatest chance of solving this impasse. Nevertheless, although it lacks a name, it would be reasonable to assume that within the sample referred to B. sp. cf. B. bandharr there is yet another late Cenozoic species of Bohra .