Tirons of the world: a review of ‘ tironid’ amphipods, description of new genera and species, and establishment of a new subfamily Tironinae Stebbing, 1906 stat. nov. (Crustacea, Synopiidae) Author Just, Jean text Zootaxa 2022 2022-05-23 5139 1 1 89 http://zoobank.org/145cb6f5-2ea8-40b0-9cca-3e942aa4a5b9 journal article 95147 10.11646/zootaxa.5139.1.1 8ab25605-ec09-4d57-9dbd-7da2b032e5e8 1175-5326 6572023 145CB6F5-2EA8-40B0-9CCA-3E942AA4A5B9 Glandulotiron intermedius ( Reid, 1951 ) comb. nov. Figs 39–40 Tiron intermedia Reid, 1951 , p. 233 , fig. 30. Tiron intermedius Reid, 1951 .—Jazdzewski 1990, (correction of epithet); Lörz & Coleman 2013 , p. 26 ; Cadien 2014 , p. 11 . Type fixation. Lectotype , ♀ .—Here designated, (the specimen partly illustrated by Reid, 1951 ). Material examined. Lectotype , ♀ , 8.0 mm. West Africa , Guinea , off Conakry , 9˚28’N, 14˚58’W, 45 m , shells, foraminifera, trawl, Sigsbee Trawl ( 1 meter ), 14. April 1946 , Atlantide stn 147, NHMD 84716 (including one slide). Paralectotypes . Same data as lectotype, NHMD 618795 ( 1 ♂ of 5.4 mm , 1 ♀ , 1 damaged ♀ , 1 juvenile ) . Description (female). Rostrum acute, apex blunt. Interantennal lobes broad, truncate with equal-sized rounded corners. Accessory eyes composed of 4 ommatidia in a curve. Mouthparts . Maxilla 1 palp with 7 subequal, pointed or minutely serrate robust setae; outer plate with row of 10 variously smooth, simple of bifurcate, pointed robust setae. Mouthparts otherwise normal. Antenna 1 as long as pereonite 1–5 combined along dorsum, peduncle article 2 about three quarter length of 1, article 3 4/5 length of article 2, article 1 with single penicillate seta ventrally in posterior half and 2 dorsally, flagellum longer than peduncle by 1/3, of 10 articles, assessory flagellum of 5+ articles. Antenna 2 as long as pereonites 1–6 and 2/3 of 7 combined along dorsum, peduncular article 4 the longest, flagellum of 8+ articles, articles 4–8 with single slender robust seta distally; otherwise setose. FIGURE 39. Glandulotiron intermedius ( Reid, 1951 ) comb. nov. Lectotype, ♀, except M , ♂, paralectotype. ant 1 and ant 2 , antenna 1 and 2; clv , head lateral view; mx1 , maxilla 1; mxp ip , maxilliped inner plate; mxp op , maxilliped outer plate. FIGURE 40. Glandulotiron intermedius ( Reid, 1951 ) comb. nov. Lectotype, ♀. cox3 , coxal plate 3; gp1–2 , gnathopods 1 and 2; p4 , pereopod 4; p5–7 , pereopods 5 to 7; tel , telson; up3 , uropod 3; us , urosome. Coxal plates 1–3 apically fringed with moderately long simple setae only, plate 4 naked, shorter than 3. Gnathopods 1 and 2 basis entire anterior margin with setae of varying length, posterior margin similar but less densely covered, carpus on both with scattered setae on anterior margin, with 10–12 groups of unequally long setae along posterior margin of carpus, propodus of gnathopod 1 with about 7 small pectinate robust setae among many long simple setae, gnathopod 2 with a few groups of setae of unequal length. Pereopods 3 and 4 sparsely setose, basis as long as remaining articles combined. Pereopods 5–6 slightly pearshaped, broadest distally, with broadly rounded posterodistal lobe, plate 5 with weakly concave posterior margin. Pereopod 7 with posteroproximal large rounded angular lobe set off from posterior margin by concavity. Pereopods 5–7 basis with row of long setae on medial surface. Uropod 1 peduncle not quite reaching end of uropod 2 peduncle, rami of equal length, 2/3 as long as peduncle, with a single small robust seta dorsally. Uropod 2 outer ramus about 4/5 length of inner ramus, reaching to about posterior margin of uropod 3 peduncle. Uropod 3 peduncle about half length of outer ramus, rami overreaching telson with about 1/5 their length, outer ramus basal article with dorsolateral row of 14–15 robust setae, ventral margin with dense felt of tiny setules, small second article with single apical robust seta, inner ramus reaching to about end of basal article of outer ramus, dorsolateral margin with row of long setae and 3 robust setae distally, apex with single robust seta. Telson . Two and half times longer than greatest width, reaching 3/4 along outer ramus of uropod 3, with row of 5 robust setae, many short and long simple setae and doublet of small plumose setae on each lobe; apex with 1 robust seta in apical hollow. Male . Generally as female but differing as follows: antenna 1 flagellum of 6 articles, article 1 with medial field of callynophore in row (possibly 2 rows) of 7 bunches of long aestetascs, articles 2–6 with single very long aestetasc (longer than in any other species in the genus), accessory flagellum of 3+ slender articles; antenna 2 flagellum of 5+ articles. Posterior expansion of basis of pereopod 7 slightly broader. Size . Largest ♂ , 5.4 mm . Largest ♀ , 8 mm . Distribution . West Africa, off Guinea . 45 m .