Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae) Author Zallot, Enrico 365E17AD-6938-4364-A526-F1BDA2663E10 Per Ligont 1, 33070 Budoia, Italy. ezallot@gmail.com Author Kamchev, Panche 9FA2BE01-3355-4340-B57C-9E2841D5E4C2 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. panekamchev@gmail.com Author Schilthuizen, Menno 683D0AB7-CDD8-4FAA-94B6-436F3BFB8873 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. & Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands. menno.schilthuizen@naturalis.nl Author Fehér, Zoltán E801EC76-8B1E-450B-993E-BBBE57C00EA9 WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary. feher.zoltan.nhmus@gmail.com Author Mattia, Willy De C74A049D-9D7B-4A1F-878B-5BDFD104219F Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria. wdemattia@gmail.com Author Gittenberger, Edmund D786C279-FC92-4D08-AF16-F79A9705E0AE Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. egittenberger@yahoo.com text European Journal of Taxonomy 2024 2024-03-21 927 1 163 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013 journal article 10.5852/ejt.2024.927.2475 2118-9773 10853688 B6E43365-FACA-49BA-8CCD-77E4BF8F0016 Cochlostoma ( T. ) nanum ( Westerlund, 1879 ) Figs 72 (red dots), 73A–C, 74–76 Pomatias nanus Westerlund, 1879: 167-16 . Pomatias clessini Hirc, 1881: 521–522 . Types Not seen. Other specimens AUSTRIA : 5- Tepke ; 46.3888° N , 14.5738° E ; 2015; AL leg.; NHMW109000 /AL/00703. SLOVENIA • 1- Mt Risnjak , 1400 m (topotypical); 45.4251° N , 14.6162° E ; 2006; De Mattia leg.; WdM5666 • 2- Mt Snežnik 1764; 45.5884° N , 14.4478° E ; 2010; De Mattia leg.; WdM7005 • 3- Mt Snežnik 1451; 45.5825° N , 14.4312° E ; 2010; De Mattia leg.; EZ0960 • 4- Logarska Dolina ; 46.4274° N , 14.6572° E ; 2009; Slapnik leg.; HNHM97239 . Type locality A generic “Croatia” in the original description. We define as type locality restricted: CROATIA • Mount Risnjak (see remarks) . Description SHELL . Closely spaced riblets on last part of protoconch. Teleoconch spotless, with weak or very weak ribs almost disappearing on last whorl and approaching aperture. Strong lip with columellar lobe either barely or abruptly inwardly bent, only partially covering umbilicus. MEASUREMENTS . 13 ♀♀ : whorls =7–8, H = 7–8.4 mm , H/W=2.4–2.64, roundness =0.1–0.15, ribs incl.= 53–61°, apert. incl.= 16–24°, ribs/mm 1 st wh.= 8–11, ribs/mm 4 th wh.=8–17. FEMALE GENITAL ORGANS . Posterior connection of pedunculus of bursa copulatrix and junction of uterus gland moved far from connection between distal oviduct and pedunculus of bursa copulatrix. Seminal receptacle normally club-shaped, situated on dorsal side of body and with long and undefined basis/ distal oviduct. Remarks The type locality in the original description is rather undefined. We consider Mount Risnjak in the northern part of Croatia as type locality restricted by A.J. Wagner (1897: 54 [618]): “Die mir vom Autor übergebenen Originalexemplare stammen ohne nähere Fundortsangabe aus Croatien (ein bestimmter Fundort wird auch sonst vom Autor nirgends angeführt); dieselben erwiesen sich aber als vollkommen identisch mit den mir ebenfalls vorliegenden Originalexemplaren des P. clessini Hirc vom Berge Risnjak in Croatien und dürften wohl auch daher stammen [The original specimens were given to me by the author labeled ‘Croatia’ without any further information on where they were found (a specific site is also not mentioned anywhere else by the author); but it turned out that they are completely identical to the original specimens of P. clessini Hirc from the Risnjak mountain in Croatia that I also have and it may well be that they are from there]”. Fig. 72. Distribution of samples of clade B: red = Cochlostoma ( T. ) nanum ( Westerlund, 1879 ) ; cyan = C. ( T. ) elegans . Black center for the samples with amplified 16S. Fig. 73. Shell and female genitals of Clade B samples with amplified 16S. A . Cochlostoma ( T. ) nanum ( Westerlund, 1879 ) , Tepke, AT (NHMW-109000/AL/00703). B . C. ( T. ) nanum , Mt Snežnik 1764 , SI (WdM-7005). C . C. ( T. ) nanum , Mt Risnjak, 1400 m a.s.l., HR (WdM-5666). D . C. ( T. ) mostarense ( Wagner, 1906 ) , Mostar, BeH (EZ-1091). E . C. ( T. ) elegans ( Clessin, 1879 ) , Razbojište near Oltari, HR (WdM-6896). F . C. ( T. ) stossichi ( Hirc, 1881 ) , 2 km NNW of Meja, HR (WdM-1240). G . C. ( T. ) braueri ( Wagner, 1897 ) , Klek Mt, HR (EZ-0068). H . NFS142, Škrčko Jezero, MONT (TxEx-du0044). I . C. ( T. ) hallgassi sp. nov. , Monte Petrella 1010, I (EZ-1118). Fig. 74. Type of Pomatias clessini Hirc, 1881 = Cochlostoma ( T. ) nanum ( Westerlund, 1879 ) , ♂, in Wagner’s collection (NHMW – without collection number) from Mount Risnjak, HR. For what is known, the species lives in a few scattered alpine localities not connected by suitable habitats. In the phylogenetic trees the nanum samples do not cluster together and the intraspecific p-distances are of the same magnitude of the ones with samples of the other species of the subclade.