Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo Author Grinang, Jongkar Author Ng, Peter K. L. text Raffles Bulletin of Zoology 2015 2015-12-18 63 564 582 journal article 10.5281/zenodo.5386711 2345-7600 5386711 BAA493D3-8195-417B-ABD8-A36D3FD90533 Lepidothelphusa loi , new species ( Figs. 7A–I , 8A–E , 14A, B ) Material examined. Holotype : male (10.3 × 9.6 mm ) ( ZRC 2015.0607 ), Gunung Ampungan, Sadong River Basin, Serian, Sarawak , coll. J Grinang et al., 8 February 2015 . Paratypes : 2 females (larger 9.8 × 9.2 mm ), 2 ovigerous females (larger 10.3 × 9.8 mm ) (SBC.C.00435–39), data same as holotype ; 14 males (largest 11.3 × 10.6 mm , 10.5 × 9.6 mm , 11.8 × 10.9 mm ), 8 females (largest 11.2 × 9.8 mm ) ( ZRC 2015.0292 ), on sides of small stream, mudstone and rocks substrate, densely forest covered slope of hill Gunung Ampungan, Sadong River Basin, Serian, Sarawak , coll. PKL Ng, J Grinang & PYC Ng, 18 and 20 June 2015 . Diagnosis. Carapace quadrate, surface relatively flat, smooth, anterolateral margins smooth; antero- and posterolateral regions rugose; epibranchial tooth undiscernible; external orbital tooth with outer margin concave, smooth; serrated projection on frontal region distinct; epigastric and postorbital cristae distinct, high, margins serrated; cervical and H-grooves relatively deep, not confluent ( Fig. 7A, B, D ); ischium of third maxilliped rhomboidal, much longer than broad, with incomplete median sulcus ( Fig. 7F ); inner margin of merus of chelipeds with relatively lower serrated projection ( Fig. 7G ); carpus of chelipeds rugose, inner angle with broad, low triangular tooth ( Fig. 7A ); fingers of adult major male chela narrowly gaping ( Fig. 7H, I ); ambulatory legs short, merus of fourth ambulatory legs about half of length of carapace ( Fig. 7A, E ); male abdomen triangular, somite 6 width and length subequal, subequal to length of telson ( Fig. 7C ); G1 stout, sinuous in ventral view, straight in dorsal view; terminal segment straight in dorsal view, gently curving outwards in ventral view ( Fig. 8A, B, D, E ); G2 with long distal segment, about three-quarters length of basal segment ( Fig. 8C ). Colour. In life, males are yellowish-red carapace, reddishbrown ambulatory legs and yellowish chelipeds. Females are light brown overall ( Fig. 14A, B ). Etymology. The name is after Michael Lo in recognition of his kind help in helping us search for these interesting animals. Remarks. Lepidothelphusa loi , new species , can be easily distinguished from L. cognettii s. str. , L. flavochela , new species , L. limau , new species , and L. padawan , new species , in having relatively shorter ambulatory legs ( Fig. 7A, E versus Figs. 1A, E , 3A, E , 5A, E , 9A, E , 11A, E ). The distal segment of the G2 of L. loi is relatively longer than congeners ( Fig. 8C versus Figs. 2C , 4C , 6C , 10C , 12C ). See also discussion for L. flavochela and L. limau . Habitat. The habitat is a gentle slope, with the substrate composed of moist mud and sand, with leaf litter. The site is shaded, has a spring, and is part of a disturbed dipterocarp forest. The base substrate is sandstone, and the site is more than 500 m above sea level.