A revision of the spirit loaches, genus Lepidocephalus (Cypriniformes, Cobitidae) Author Deein, Gridsada Inland Fisheries Research and Development Bureau, Department of Fisheries, NIFI Building, Kasetklang, Chatuchak, Bangkok 10900 Thailand. E-mail: dgridsada @ hotmail. com Author Tangjitjaroen, Weerapongse Faculty of Veterinary Medicine, Chiang Mai University, Chiang Mai, 50100, Thailand. E-mail: weerapongse @ yahoo. com Author Page, Lawrence M. Florida Museum of Natural History, University of Florida, Gainesville, FL 32611 - 7800 USA. E-mail: lpage 1 @ ufl. edu text Zootaxa 2014 2014-03-17 3779 3 341 352 journal article 5815 10.11646/zootaxa.3779.3.2 2055f347-d478-4d50-aad0-7e78c4b3b9c5 1175-5326 4910403 23584B50-EFAC-4BCE-A20E-9B09C22529B0 Key to species of Lepidocephalus 1. Eyes and dark pigment absent; ventrally directed tubules along lateral line; western Kalimantan .............. L. spectrum 1. Eyes and dark pigment present; no tubules along lateral line.................................................... 2 2. Dorsal-fin origin over pelvic-fin origin; western Kalimantan ........................................... L. pallens 2. Dorsal-fin origin well behind pelvic-fin origin............................................................... 3 3. No scales on top of head; 9 branched dorsal-fin rays; 4 branched anal-fin rays; peninsular Malaysia ........ L. pahangensis 3. Scales (sometimes embedded) on top of head; 8 branched dorsal-fin rays; 5–6 branched anal-fin rays.................. 4 4. Snout short, 3.9–5.1% SL, 23.3–27.1% HL; Java, Sumatra , southern Kalimantan ......................... L. macrochir 4. Snout longer, 5.6–7.4% SL, 28.6–38.1% HL; Thailand ................................................ L. nanensis