The millipede family Paradoxosomatidae in the Philippines, with a description of Eustrongylosomapenevi sp. n., and notes on Anoplodesmusanthracinus Pocock, 1895, recorded in Malaysia and Sri Lanka for the first time (Diplopoda, Polydesmida) Author Golovatch, Sergei Author Stoev, Pavel text Biodiversity Data Journal 2013 1 957 957 http://dx.doi.org/10.3897/BDJ.1.e957 journal article http://dx.doi.org/10.3897/BDJ.1.e957 1314-2828-1-957 Rank: SpeciesType of treatment: New taxonextantHabitat: terrestrialRoot classification: 8 Eustrongylosoma penevi Golovatch & Stoev, 2013 sp. n. Materials Type status: Holotype Occurrence: recordedBy: P. Stoev & L. Penev ; individualCount: 1 ; sex: male ; Location: island: Luzon Island; country: Philippines ; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation: 1800-1900 m; locationRemarks: under stones and logs; verbatimLatitude: 16°57\'58"N; verbatimLongitude: 121°1\'37"E; Event: eventDate: 6 July 2012 ; Record Level: institutionCode: NMNHS Type status: Paratype Occurrence: recordedBy: P. Stoev & L. Penev ; individualCount: 4 ; sex: male ; Location: island: Luzon Island; country: Philippines ; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation: 1800-1900 m; locationRemarks: under stones and logs; verbatimLatitude: 16°57\'58"N; verbatimLongitude: 121°1\'37"E; Event: eventDate: 6 July 2012 ; Record Level: institutionCode: NMNHS Type status: Paratype Occurrence: recordedBy: P. Stoev & L. Penev ; individualCount: 1 ; sex: female ; Location: island: Luzon Island; country: Philippines ; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation: 1800-1900 m; locationRemarks: under stones and logs; verbatimLatitude: 16°57\'58"N; verbatimLongitude: 121°1\'37"E; Event: eventDate: 6 July 2012 ; Record Level: institutionCode: NMNHS Type status: Paratype Occurrence: recordedBy: P. Stoev & L. Penev ; individualCount: 1 ; lifeStage: juvenile ; Location: island: Luzon Island; country: Philippines ; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation: 1800-1900 m; locationRemarks: under stones and logs; verbatimLatitude: 16°57\'58"N; verbatimLongitude: 121°1\'37"E; Event: eventDate: 6 July 2012 ; Record Level: institutionCode: NMNHS Type status: Paratype Occurrence: recordedBy: P. Stoev & L. Penev ; individualCount: 1 ; sex: male ; Location: island: Luzon Island; country: Philippines ; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation: 1800-1900 m; locationRemarks: under stones and logs; verbatimLatitude: 16°57\'58"N; verbatimLongitude: 121°1\'37"E; Event: eventDate: 6 July 2012 ; Record Level: institutionCode: ZMUM Description Length 18-22 (♂) or 23 mm (♀), width of midbody pro- and metazona 1.1-1.3 and 1.5-1.7 mm (♂), or 1.9 and 2.1 mm (♀), respectively. Holotype ca 22 mm long, width of pro- and metazona 1.3 and 1.6 mm, respectively. Coloration black to light grey-brown (Fig. 1a). Pattern mostly cingulate due to a large light grey band on prozona dorsally in front of stricture extending down until level of paraterga (Fig. 1a, b). Legs light grey-brown. Antennae increasingly infuscate distad, from light brown to blackish (Fig. 1a). Body submoniliform. Antennomeres 2 to 6 subequal in length, antennae rather short, reaching behind segment 3 (♂) or 2 (♀) when stretched dorsally. Tegument generally smooth and shining. In width, segments 2 and 3 <collum = 4 <5-17, thereafter body gently tapering towards telson. Paraterga (Fig. 1b, c, d) modestly developed, keel-shaped, set low (at about 1/3rd of metazonal height), thinner in poreless, thicker in pore-bearing, segments, slightly reaching behind tergal margin only in segments 2 and 17-19, mostly slightly pointed, delimited by a complete and deep sulcus only dorsally, ventral sulcus being incomplete, developed only in posterior quarter to 1/5th extent. Head densely setose on clypeus and frons, bare on vertex. Collum semilunar, bearing two transverse rows of 2+2 setae, one row along front margin, second row in the middle; lateral edges broadly rounded. Metaterga faintly rugulose, a little more clearly so in postsulcus halves, surface below paraterga microgranular in segments 2-7. Tergal setae rather long, about 1/5th of metatergal length, arranged in two rows of 2+2 in each, one in front of, second behind sulcus. Sulcus starting from segment 5, deep, almost reaching the bases of paraterga. Stricture between pro- and metazona finely and densely ribbed. Ozopores lying close to caudal end of paraterga in a shallow ovoid groove, lateral, only partly visible from above. Pleurosternal carinae poorly developed ridges visible only in segments 2-4. Seta at about midway of each paratergum mostly broken off. Axial line wanting. Epiproct (Fig. 2a) subtruncate, pre-apical lateral papillae small. Hypoproct (Fig. 2b) semi-circular. Sternal lobe between coxae 4 subtrapeziform, densely setose (Fig. 2c). Legs very long and slender, about 2.0 (♂) or 1.3 (♀) times as long as midbody height; ♂ tarsal brushes traceable until about legs of segment 15, thereafter thinning out. Gonopods rather simple (Fig. 2d, e, f): coxite long, subcylindrical, bare; prefemoral portion small, about 1/3rd as long as femorite, the latter slender, ventral lobe somewhat better developed than dorsal one, apicolateral lobe (l) rounded, well developed, with a long transverse spine (s) at base. Solenophore subcircular, with a subterminal lobule. Diagnosis Most similar to Eustrongylosoma exiguum Hoffman, 1978, from Papua New Guinea, and Eustrongylosoma kuekenthali (Attems, 1897), from Borneo and Sulawesi, sharing the presence of a prominent distal spine on the gonopod femorite. Different from all congeners by the transverse orientation of the spine and noticeably long legs in the male ( Hoffman 1978 , Golovatch 1997 ). Etymology Honours our good friend and colleague Lyubomir Penev, biologist and founder of the Biodiversity Data Journal and Pensoft Publishers. <br/> Notes The species is hitherto known only from its type locality, Mt Polis Checkpoint on the road Banaue - Sagada (Fig. 3), where it was found close to a human settlement, under wooden plates and logs (Fig. 4).