Review of Cycadophila Xu, Tang & Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species Author Skelley, Paul Author Xu, Guang Author Tang, William Author Lindström, Anders J. Author Marler, Thomas Author Khuraijam, Jibankumar Singh Author Singh, Rita Author Rich, Stephen text Zootaxa 2017 4267 1 1 63 journal article 33061 10.5281/zenodo.575641 810b55e4-b8ba-4368-bf9c-8517ee33e7d0 1175-5326 575641 8920893E-E5F8-482A-A60D-7A248D2B0DCB Cycadophila ( Strobilophila ) 16(1). Protibia notably dilated toward apex with stout marginal spines on lateral apical 1/3 separated by less than their length ( Figs. 2 F, 20D); with distinct furrow on each side of pronotal base ( Fig. 19 A)........................................ 17 - Protibia weakly dilated distally with stout marginal spines on lateral apical 1/2 separated by their length or more ( Figs. 2 G, 21D); lacking or with greatly reduced furrow on each side of pronotal base ( Fig. 21 A); Thailand .............................................................................................. Cycadophila ( S. ) yangi , new species 17(16). Submental-gular suture indicated by weakly arcuate row of punctures; basal margin of pronotum nearly straight from side to side, weakly convex, lacking weak angular concavities where pronotal discal grooves join margin; scutellar shield more pentagonal; female with terminal abdominal ventrite convex apically, unmodified ( Figs. 16 G, 19G); male flagellum and penile struts longer, length of median lobe to penile strut 1:5; Thailand, India.......................................... 18 - Submental-gular suture indicated by distinctly arcuate row of punctures; basal margin of pronotum sinuate, with distinct but shallow angular concavities where pronotal discal grooves join margin; scutellar shield more transverse; female with terminal abdominal ventrite concave apically, modified ( Fig. 17 E); male flagellum and penile struts shorter, length of median lobe to penile strut 2:7; Vietnam ............................................................ C. ( S. ) hiepi , new species 18(17). Posterior lateral margin of pronotum straight to posterior angle which may have a very small indistinct projection behind pore ( Fig. 18 A); no sexual dimorphism visible on terminal abdominal ventrite; inhabiting Cycas sp. on karst towers near the city of Kanchanaburi , Kanchanaburi province in Thailand at elevations below 100 m . ............ C. ( S. ) kwaiensis , new species - Posterior lateral margin of pronotum distinctly curved inward at posterior angle which has a distinct, small acute projection behind pore ( Figs. 16 A,D, 19A); females with terminal abdominal ventrite with a small area of fine punctures on apical medial surface, males evenly coarsely punctate throughout; occurring in widespread localities and hosts..................... 19 19(18). Elytral striae weakly impressed, giving intervals a weakly convex appearance, most evident on middle of disc; female terminal abdominal ventrite with apical medial surface glossy, glabrous and finely punctate, distinctly different from rest of surface, males with this surface entirely setose and coarsely punctate ( Figs. 19 F–G) similar to rest of surface; inhabiting Cycas elephantipes , C. clivicola lutea , C. pectinata , C. petraea and C. tansachai above 300 m elevation in Thailand ............................................................................................ C. ( S. ) tansachai , new species - Elytral striae not impressed, intervals appearing flat; both male and female terminal abdominal ventrite with apical medial surface setose, females with finer punctures on apical medial surface than males ( Figs. 16 F–G); inhabiting Cycas pectinata , Assam , India ................................................................ C. ( S. ) assamensis , new species