<document id="613642835BB1BBE8956C3DE15D1B866E" ID-CLB-Dataset="31855" ID-DOI="10.11646/zootaxa.4331.1.1" ID-GBIF-Dataset="14c29731-224f-43a4-ae52-f76823177ce5" ID-ISSN="1175-5326" ID-Zenodo-Dep="1009725" ID-ZooBank="FB50725E-4D03-4507-9CAF-EB939C7F548F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507876567978" checkinUser="plazi" docAuthor="Asami, Takahiro" docDate="2017" docId="03CA87D8FFB5FF9EFF0BFF58FD54FE1C" docLanguage="en" docName="zootaxa.4331.1.1.pdf" docOrigin="Zootaxa 4331 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Metalycaeus Pilsbry 1900" docType="treatment" docVersion="9" lastPageNumber="73" masterDocId="FFF3FFA0FFFDFFD6FF9CFFC0FF80FFFF" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus" masterLastPageNumber="124" masterPageNumber="1" pageNumber="73" updateTime="1698469440756" updateUser="ExternalLinkService">
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<mods:title id="8F07F1B2842E4954EAE4363819BFC7F3">Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus</mods:title>
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<mods:namePart id="D75C5EB0CC1240EAE8DC1396BAB03C4D">Asami, Takahiro</mods:namePart>
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<mods:date id="8D4FA9545FC2A41342262447F9C4C4D2">2017</mods:date>
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<treatment id="03CA87D8FFB5FF9EFF0BFF58FD54FE1C" ID-DOI="http://doi.org/10.5281/zenodo.6004460" ID-GBIF-Taxon="135435113" ID-Zenodo-Dep="6004460" LSID="urn:lsid:plazi:treatment:03CA87D8FFB5FF9EFF0BFF58FD54FE1C" httpUri="http://treatment.plazi.org/id/03CA87D8FFB5FF9EFF0BFF58FD54FE1C" lastPageNumber="73" pageId="72" pageNumber="73">
<subSubSection id="C3796545FFB5FF9EFF0BFF58FDAEFF4D" box="[151,558,151,178]" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="8BDC36CEFFB5FF9EFF0BFF58FDAEFF4D" blockId="72.[151,558,151,178]" box="[151,558,151,178]" pageId="72" pageNumber="73">
<heading id="D09481A2FFB5FF9EFF0BFF58FDAEFF4D" bold="true" box="[151,558,151,178]" fontSize="11" level="1" pageId="72" pageNumber="73" reason="1">
<emphasis id="B917EADCFFB5FF9EFF0BFF58FDAEFF4D" bold="true" box="[151,558,151,178]" pageId="72" pageNumber="73">
Genus 
<taxonomicName id="4C634D4DFFB5FF9EFF72FF57FDAEFF4D" ID-CoL="7P8SD" authority="Pilsbry, 1900" authorityName="Pilsbry" authorityYear="1900" box="[238,558,151,178]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="73" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFB5FF9EFF72FF57FE05FF4E" bold="true" box="[238,389,151,177]" italics="true" pageId="72" pageNumber="73">Metalycaeus</emphasis>
Pilsbry, 1900
</taxonomicName>
</emphasis>
</heading>
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<subSubSection id="C3796545FFB5FF9EFF0BFF19FE56FF10" box="[151,470,217,239]" pageId="72" pageNumber="73" type="reference_group">
<paragraph id="8BDC36CEFFB5FF9EFF0BFF19FE56FF10" blockId="72.[151,470,217,239]" box="[151,470,217,239]" pageId="72" pageNumber="73">
<taxonomicName id="4C634D4DFFB5FF9EFF0BFF19FE52FF10" authority="Pilsbry, 1900: 382" authorityName="Pilsbry" authorityPageNumber="382" authorityYear="1900" box="[151,466,217,239]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="73" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFB5FF9EFF0BFF19FE90FF10" box="[151,272,217,239]" italics="true" pageId="72" pageNumber="73">Metalycaeus</emphasis>
Pilsbry, 1900: 382
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFB5FF9EFF0BFED6FB8DFEAC" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="8BDC36CEFFB5FF9EFF0BFED6FB8DFEAC" blockId="72.[151,1437,278,483]" pageId="72" pageNumber="73">
<emphasis id="B917EADCFFB5FF9EFF0BFED6FEBBFED0" bold="true" box="[151,315,278,303]" pageId="72" pageNumber="73">Type species.</emphasis>
<taxonomicName id="4C634D4DFFB5FF9EFED6FED6FC2BFED0" authority="PILsbry, 1900" authorityName="PILsbry" authorityYear="1900" box="[330,939,278,303]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="72" phylum="Mollusca" rank="species" species="melanopoma" subGenus="Metalycaeus">
<emphasis id="B917EADCFFB5FF9EFED6FED6FE2FFED1" box="[330,431,278,302]" italics="true" pageId="72" pageNumber="73">Alycaeus</emphasis>
(
<emphasis id="B917EADCFFB5FF9EFE54FED6FDD5FED1" box="[456,597,278,302]" italics="true" pageId="72" pageNumber="73">Metalycaeus</emphasis>
) 
<emphasis id="B917EADCFFB5FF9EFDF3FED6FC80FED1" box="[623,768,278,302]" italics="true" pageId="72" pageNumber="73">melanopoma</emphasis>
PILsbry, 1900
</taxonomicName>
(syNONym Of 
<taxonomicName id="4C634D4DFFB5FF9EFBFEFED6FA1CFED1" box="[1122,1436,278,302]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="nipponensis">
<emphasis id="B917EADCFFB5FF9EFBFEFED6FA1CFED1" box="[1122,1436,278,302]" italics="true" pageId="72" pageNumber="73">Chamalycaeus nipponensis</emphasis>
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[REINHardT, 1877], sEE 
<bibRefCitation id="EFF24B3FFFB5FF9EFE0BFEFBFDA9FEAC" author="MINaTO" box="[407,553,315,339]" pageId="72" pageNumber="73" refString="Minato, H. (1988) A systematic and bibliographic list of the Japanese land snails. Hiroshi Minato, Shirahama, 294 pp." type="book" year="1988">MINaTO 1988</bibRefCitation>
), by sUbsEqUENT dEsIGNaTION (
<bibRefCitation id="EFF24B3FFFB5FF9EFCEEFEFBFB80FEAC" author="THIELE" box="[882,1024,315,339]" pageId="72" pageNumber="73" refString="Thiele J., (1929) Handbuch der systematischen Weichtierkunde. Teil 1. Verlag von Gustav Fischer, Jena, 376 pp." type="book" year="1929">THIELE 1929</bibRefCitation>
).
</paragraph>
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<subSubSection id="C3796545FFB5FF9EFF5BFE9EFD54FE1C" pageId="72" pageNumber="73" type="discussion">
<paragraph id="8BDC36CEFFB5FF9EFF5BFE9EFD54FE1C" blockId="72.[151,1437,278,483]" pageId="72" pageNumber="73">
<emphasis id="B917EADCFFB5FF9EFF5BFE9EFEBBFE88" bold="true" box="[199,315,350,375]" pageId="72" pageNumber="73">Remarks.</emphasis>
THE GENErIC LEvEL rEvIsION Of THE 
<taxonomicName id="4C634D4DFFB5FF9EFD2DFE9EFCC9FE89" authority="Is" authorityName="Is" box="[689,841,350,375]" class="Gastropoda" family="Alycaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="73" phylum="Mollusca" rank="family">ALyCaEIdaE Is</taxonomicName>
IN prOGrEss. HErE wE rEfraIN frOm GIvING a dIaGNOsIs, aNd TENTaTIvELy CLassIfy aLL CHINEsE, LaO aNd VIETNamEsE spECIEs IN 
<taxonomicName id="4C634D4DFFB5FF9EFC25FE43FBC6FE64" authorityName="Pilsbry" authorityYear="1900" box="[953,1094,387,411]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="73" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFB5FF9EFC25FE43FBC6FE64" box="[953,1094,387,411]" italics="true" pageId="72" pageNumber="73">Metalycaeus</emphasis>
</taxonomicName>
, wHICH HavE spIraLLy sTrIaTEd prOTOCONCHs. BOTH 
<taxonomicName id="4C634D4DFFB5FF9EFEF2FE66FE7CFE41" authorityName="Pilsbry" authorityYear="1900" box="[366,508,422,446]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="73" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFB5FF9EFEF2FE66FE7CFE41" box="[366,508,422,446]" italics="true" pageId="72" pageNumber="73">Metalycaeus</emphasis>
</taxonomicName>
aNd 
<taxonomicName id="4C634D4DFFB5FF9EFDAAFE66FD5BFE41" authorityName="Mollendorff" authorityYear="1897" box="[566,731,422,446]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="72" pageNumber="73" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFB5FF9EFDAAFE66FD5BFE41" box="[566,731,422,446]" italics="true" pageId="72" pageNumber="73">Chamalycaeus</emphasis>
</taxonomicName>
HavE spIraL sTrIaE ON THE TELEOCONCH, bUT THE spIraL sTrIaTION Is mIssING ON THE prOTOCONCH Of THE LaTTEr (
<figureCitation id="13582A4BFFB5FF9EFDC8FE0BFD48FE1C" box="[596,712,458,483]" captionStart="FIGURE 46" captionStartId="71.[151,250,1777,1799]" captionTargetBox="[222,1373,193,1762]" captionTargetId="figure@71.[222,1373,193,1762]" captionTargetPageId="71" captionText="FIGURE 46. Type species of alycaeid genera and SEM images of their embryonic whorls. A – C. Metalycaeus nipponensis (Reinhardt, 1877). A. lectotype (ANSP 78815) of Alycaeus (Metalycaeus) melanopoma Pilsbry, 1900, the type species of Metalycaeus Pilsbry, 1900, Hakone Mts., Japan, leg. Schmacker. B – C. Japan, Stukui Town, Shida pass, leg. K. Ohara, 0 6 June 1986, HNHM 99701. D – F. Alycaeus (Chamalycaeus) fruhstorferi Möllendorff, 1897 (type species of Chamalycaeus Möllendorff, 1897). D. Java, leg. Fruhstorfer, coll. Möllendorff (lectotype, SMF 109481). E – F. W. Java, Mt. Gede, 4000 ', leg. Fruhstorfer, Aug 1892, E. R. Sykes coll. 1954, NHMUK 20150361. Photos: Anonymous (A: downloaded from the website of the ANSP); B. Páll-Gergely (B – C, E – F); F. Walther (D). Small white arrows indicate the protoconch-teleoconch boundary. Uppermost scale represents 5 mm and refers to figs A and D." httpUri="https://zenodo.org/record/1009817/files/figure.png" pageId="72" pageNumber="73">FIGUrE 46</figureCitation>
).
</paragraph>
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</treatment>
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