Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa Author GUINOT, DANIÈLE Author TAVARES, MARCOS Author CASTRO, PETER text Zootaxa 2013 2013-05-29 3665 1 1 414 http://dx.doi.org/10.11646/zootaxa.3665.1.1 journal article 10.11646/zootaxa.3665.1.1 1175-5326 5276390 8358B363-BEE3-416D-96CA-8614E38B61D5 Family † Telamonocarcinidae Larghi, 2004 new status Onomatophore (= type genus). † Telamonocarcinus Larghi, 2004 (Cenomanian) . Genera (tentatively) included. † Telamonocarcinus , with † T. gambalatus Larghi, 2004 , as type species; † Eodorippe Glaessner, 1980 , with † E. spedeni Glaessner, 1980 (Campanian–Maastrichtian) as type species; † Tepexicarcinus Feldmann, Vega, Applegate & Bishop, 1998 , with † T. tlayuaensis Feldmann, Vega, Applegate & Bishop, 1998 (Albian) as type species. Diagnosis . Carapace from narrow, slightly pyriform to subrectangular in outline; dorsal surface well marked by several grooves; meso- and urogastric lobes well-defined. Anterolateral margin unarmed or with 1 or 2 teeth posterior to exorbital teeth. Eyes large, with long eyestalks. Chelipeds subequal. P2, P3 well developed, elongated, with wide meri, carpi and long, curved dactyli. P4, P5 reduced, probably without subchelate dactyli. Thoracic sternum pentagonal, with large sternites 5, 6; sternal sutures 4/5–6/7 incomplete. Absence of female gonopores on P3 coxae. Male abdomen with first 3 somites dorsally exposed, composed of 6 free somites plus telson († Telamonocarcinus ), 5 somites plus telson († Tepexicarcinus ) , or unknown († Eodorippe ). Remarks. † Telamonocarcininae Larghi, 2004 , is elevated at a familial rank within Dorippoidea (see Fossil Dorippoidea ). The “long and narrow, spatulate, pointing forward rostrum” of † Eodorippe spedeni Glaessner, 1980 ( Glaessner 1980: 183 , figs. 13, 13a), could correspond to the endostomial gutter that projects in the median frontal incision in extant dorippids and is visible dorsally, but is never as developed as figured for † E. spedeni . The absence of P5 subchelate dactyli is as in Ethusidae . The male abdomen consists of six free somites plus telson as in extant Dorippidae , or of five somites plus telson as in extant Ethusidae . The thoracic sternum is similar to that of dorippoids.