A new genus and eight new species of tail-wagger snails from eastern South Africa, with a key to genera within Sheldonia s. l. (Gastropoda: Urocyclidae) Author Herbert, David G. text European Journal of Taxonomy 2017 2017-04-10 309 1 50 journal article 22144 10.5852/ejt.2017.309 4e7bcd4d-1cdb-45cf-b2c3-54da2466f171 2118-9773 3827189 1E8FE779-D6E7-428E-9538-5E5F8ECFB271 Key to genera within Sheldonia s.l . 1. Epiphallic caecum arising close to penis apex and adjacent to insertion of penial retractor muscle ( Fig. 1A ) ……………………………………………………………………………………………2 – Epiphallic caecum arising nearer to mid-point of epiphallus ( Fig. 1 B–C) …………………………3 2. Flagellum short and stout ( Fig. 1D ); epiphallus short; vas deferens thick, long and convoluted; spermatophore tail plumose ( Fig. 1E );protoconch spirally punctate ……… Ptilototheca Herbert, 2016 – Flagellum longer, divided into distinct f1 and f2 components ( Fig. 1A ); vas deferens slender and relatively short, not convoluted; spermatophore tail with branched spines ( Fig. 1F ), but not plumose; protoconch smooth (rarely spirally lirate), not punctate. ……… Kerkophorus Godwin-Austen, 1912 3. Epiphallic caecum small and globular ( Fig. 1B ); spermatophore tail bifid ( Fig. 1G ) ……………… ………………………………………………………………… Microkerkus Godwin-Austen, 1912 – Epiphallic caecum large, cylindrical and usually curved ( Fig. 1C ); spermatophore tail not bifid …4 4. Protoconch spirally punctate or with numerous, close-set, incised spiral microstriae; nonspinose distal portion of spermatophore tail long, the subterminal region with short, stout projections on both sides ( Fig. 1H , inset) …………………………… Sheldonia s.s. Ancey, 1887 – Protoconch essentially smooth; non-spinose distal portion of spermatophore tail short, lacking lateral projections ( Fig. 1I , arrow) ………………………………………… Selatodryas gen. nov. This key must be considered provisional and is provided as a guide to the identification of the existing supra-specific taxa within Sheldonia s.l. However, there remain taxa within Sheldonia s.l . that do not fit well into any of these taxa (e.g., Sheldonia monsmaripi Herbert, 2016 and S. wolkbergensis Herbert, 2016 ). Establishing the true relationships of these enigmatic species will require additional insights from molecular data. Furthermore, even within these genera, there is significant variation and future research may reveal that this encompasses additional lineages worthy of recognition as distinct genera, particularly within Kerkophorus and Microkerkus . Construction of a key should thus be viewed as an iterative process, for which this is a first step.