Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae)
Author
Pekarsky, Oleg
H- 1068, Budapest, Felsoerdosor u. 16 - 18, Hungary
opbp@t-online.hu
text
ZooKeys
2014
2014-11-05
452
107
129
http://dx.doi.org/10.3897/zookeys.452.8152
journal article
http://dx.doi.org/10.3897/zookeys.452.8152
1313-2970-452-107
F7F5E4C32D764ACAA4C35B8DE5D0F90C
FFDEFFD61800FFEEBA2CFFA1014FFF8A
578587
Lygephila lupina (Graeser, 1890)
stat. n.
Figs 64
, 65
Toxocampa
lupina
Graeser, 1890, Berliner Entomologische Zeitschrift, 35: 71-84. (TL: [Russia, Judish Autonomy, Radde] Raddefka)
Lygephila mirabilis
(Bryk, 1948),
syn. n.
Eccrita mirabilis
Bryk, 1948 (TL: N Korea, Shuotsu)
Type material examined.
♀ Type, Amur Centr. (Radde), [18]87, ex. coll.
Puengeler
, slide No. OP1931f (coll. MNHU).
Additional
material examined.
1 ♀, [RFE], Ussuriysk dist., Kajmanovka, 20.VII.[19]82, leg. Ivanov, slide No. 0321Matov (coll. ZISP). 1 ♀, [China], Tapaishan im Tsinling, Sued-Shensi, Ca. 1700 m, 14.7.1936, H.
Hoene
, slide No. OP2402f, 1 ♀, [China], Tapaishan im Tsinling, Sued-Shensi, Ca. 1700 m, 10.8.1936, H.
Hoene
, slide No. OP2403f (coll. ZFMK).
Note.
There is a lot of confusion between
Lygephila mirabilis
and
Lygephila vulcanea
(Butler, 1881) in the literature with regard to illustrations of the adults and genitalia. The taxonomic clarification of this problem will be given in a separate publication.
Taxonomy.
The identity of
Lygephila lupina
was unclear for a long time.
Lygephila lupina
was described, according to the original description, from Radde, central Amur [Khabarovsk region] (Graeser 1890) on the basis of a single male from the collection of Taylor
Tancre
, in comparison with
Lygephila maxima
(Bremer, 1861). The
Puengeler
collection, now in MNHU, contains a female specimen with a type label (Fig.
64
). One can be convinced from the information given by the handwriting of
Puengeler
on the opposite side of the collecting label (made in May 1905) that this is a true type specimen from the
Tancre
collection and it is a female, not a male. Thus, Graeser was mistaken about the sex of the type specimen. The study of the genitalia of the type specimen reveals that
Lygephila lupina
is conspecific with the species known as
Lygephila mirabilis
(Bryk, 1948).
Lygephila mirabilis
, therefore, is a junior synonym of
Lygephila lupina
, syn. n.
Figures 64-69.
64-66
Adults. 64, 65
Lygephila lupina
(=
mirabilis
)
64
♀, Type, Russia, Raddefka
65
Russia, Kajmanovka
66
Lygephila vulcanea
♀, Japan
67-69
Female genitalia
67, 68
Lygephila lupina
(=
mirabilis
) 67 Russia, Raddefka, slide No. OP1931f
68
Russia, Kajmanovka, slide No. 0321Matov
69
Lygephila vulcanea
, Japan, slide No. OP2442f.
Diagnosis.
The distinctive features of
Lygephila lupina
and
Lygephila vulcanea
(Fig.
66
) are given in the works of
Sviridov (1990)
and
Kononenko (1996)
(under the names
Lygephila vulcanea
and
Lygephila mirabilis
). The main external differences between the two species are found in the colouration and shape of the forewing:
Lygephila lupina
is broader winged and paler in colouration, usually ochreous brown to buff coloured, whereas
Lygephila vulcanea
is darker, deep brown to claret brown, most often with a clearly visible violaceous shade and the forewing apex is somewhat more pointed. In the majority of the specimens the reniform stigma of
Lygephila lupina
is stronger, sharper, and more distinctly marked against the paler background. The female genitalia differ from those of
Lygephila vulcanea
(Fig.
69
) by the shallower incision of the posterior margin of the antrum.
Description.
Wingspan 44-49 mm. Head and body brownish grey; collar dark chocolate brown. Forewing brownish grey with sparse dark brown irroration; subbasal line indistinct; antemedial line arched with costal patch; reniform stigma large, dark brown, consists of 5 or 6 streak-like spots; orbicular stigma absent; postmedial and subterminal lines distinct; terminal line a row of black dots. Hindwing brownish; small discal spot present on underside; outer third dark brown; fringes as ground color.
Female genitalia
(Figs
67
,
68
). Ovipositor long, papillae anales large, hairy with long setae on apical edges. Apophyses anteriores relatively short, apophyses posteriores thin, longer than apophyses anteriores. Antrum long, narrow anteriorly, dilated posteriorly, ostium broad, posterior margin with small U-shaped cleft. Corpus bursae membranous, ovoid.
Distribution.
Russian Far East, China, Korea.