Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species
Author
Renan, Ittai
Author
Assmann, Thorsten
Author
Freidberg, Amnon
text
ZooKeys
2018
753
23
82
http://dx.doi.org/10.3897/zookeys.753.22366
journal article
http://dx.doi.org/10.3897/zookeys.753.22366
1313-2970-753-23
BBC59C6140DD44F3B9F34C011E0D0B75
BBC59C6140DD44F3B9F34C011E0D0B75
Graphipterus reymondi Antoine, 1953: 208
stat. n.
Figs 6b, 9k, 16, 25b
Types.
Holotype: ♂ (White label, brown handwritten): <Reymondi m./ (same label, black typewritten): Antoine det.>. (White label, black handwritten): <Inhamid/ Sahara septe./(Reymond)>. (Red label, black handwritten): <HOLOTYPE>. Deposited in NHMB, General collection, box 31[examined].
Diagnosis.
Large species with 20-24 isolated white round spots on elytra, six marginal extensions, extension II short, almost triangular. Humeri very narrowed, maximum width of elytra at interior rear third. The discal elytra pattern comprises a group of 8-12 elongated spots in an order parallel to the suture. Median lobe of aedeagus with ventrally bent tip.
Comparisons.
Graphipterus reymondi
resembles
G. sharonae
sp. n., from which it differs mainly by mentum and humeri morphology, pattern, color and morphology of elytra (see full comparisons under
G. sharonae
sp. n.).
Graphipterus reymondi
resembles also
G. stagonopsis
sp. n., from which it differs mainly by mentum morphology, pattern, and morphology of elytra, and color of claws and spurs (see full comparisons under
G. stagonopsis
sp. n.).
Description.
BL male: 17-18, average 17.6
+/-
0.4 mm; BL female: 17.4-21.4, average 19.3
+/-
2 mm.
Head medium; HW/PW: 0.76; EYL: 1.6-1.8 mm; EYL/EL: 0.17. Mentum with three teeth (cf. Fig. 3d). Frontal ridge absent. In male, apical white frons stripes wider than exposed frons (Fig. 4b).
Pronotum wide; PL/PW: 0.72; BPW/PW: 0.63; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long.
Elytra with strongly narrowed humeri; EL: 9.4-10.3 mm, average 9.75 mm; EW: 8.0-8.5, average 8.3 mm; EL/EW: 1.2. Lateral cross section convex. Suture conspicuous. Scales brown, disc visible between them (Fig. 6b). White lateral margin nearly as wide as antennomere I long and with six extensions; extension I triangular with rounded tip, slightly more elongated than in
G. serrator
, wider and shorter than extensionII; the latter one elongated, at third quarter of elytra. White posterior margin commonly slightly wider than lateral margin, gap at suture smaller than lateral margin, usually with a small, indistinct tip anteriorly. Disc usually with 20-24 (rarely 18), mainly rounded spots; anterior pair of spots rounded, wide as extension I, usually smaller than posterior spots, but larger than spots on central disc; central disc spots usually asymmetrically smeared; posterior pair of spots rounded; one or two small additional spots adjacent laterally to the posterior ones. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (Fig. 7a).
Legs long; MTIL: 5.8-6.1 mm, average 5.9 mm; El/MTIL: 1.6. Metatibial secondary spur brown. MTAL length: 4.2-5, average 4.7 mm; MTAL/MTIL: 0.8. Claws of hind legs black at base.
Median lobe of aedeagus with ventrally bent tip (Fig. 9k).
Habitat.
Unknown.
Co-occurring species.
No co-occurring species.
Distribution.
Morocco (Fig. 16).
Conservation.
Unknown.