<document id="8192C336DBA817731F0E7FA95B1F6EF1" ID-DOI="10.5281/zenodo.5353792" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353792" ID-ZooBank="88D45C5E-0E49-4F02-87A4-C4FA55A83B2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212659467" checkinUser="felipe" docAuthor="Freese, Edwin L., Veal, Doug A. &amp; Lago, Paul K." docDate="2020" docId="28608792FF92366EFF76F96AB92DFF6B" docLanguage="en" docName="0787.pdf" docOrigin="Insecta Mundi 2020 (787)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Liothorax alternatus" docType="treatment" docVersion="3" lastPageNumber="20" masterDocId="D459FFEAFF86367BFFB6FFEDBB37FF84" masterDocTitle="The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist" masterLastPageNumber="83" masterPageNumber="1" pageNumber="19" updateTime="1699188000474" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="7A6EF3CCEFE7634F046903419E1222FB">The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist</mods:title>
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<mods:namePart id="D283B38A5BB0613090BE40652077CF67">Freese, Edwin L.</mods:namePart>
<mods:affiliation id="48FFD65A8FE153D38F92D9C779E9887E">33493 “ S ” Avenue Adel, Iowa 50003</mods:affiliation>
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<mods:namePart id="1B233F6EF3B13B9851702BCF8B052951">Veal, Doug A.</mods:namePart>
<mods:affiliation id="F3DADBE3E0FBB8A04F8DC286DE0A6004">168 Thompson Drive SE Cedar Rapids, Iowa 52403</mods:affiliation>
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<mods:namePart id="EDAB5C0FBD52600093B15BE723B52125">Lago, Paul K.</mods:namePart>
<mods:affiliation id="6B696058922A73EBFB223139AF7FA09C">Biology Department, University of Mississippi University, Mississippi 38677</mods:affiliation>
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<mods:title id="454D4A01FFED48965C576EB108334859">Insecta Mundi</mods:title>
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<mods:date id="83EFD0E53958AC14497760A44163A1B7">2020</mods:date>
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<mods:number id="7BC7445A2E0CB9C4227DC76B43B5A5A1">787</mods:number>
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<mods:identifier id="29B3F415B9597D73E542885629AFC45A" type="ISSN">1942-1354</mods:identifier>
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<taxonomicName id="67C94D07FF92366FFF76F96AB9B9F91B" authority="(Horn, 1870)" baseAuthorityName="Horn" baseAuthorityYear="1870" box="[192,654,1671,1695]" class="Insecta" family="Aphodiidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="alternatus">
<emphasis id="92BDEA96FF92366FFF76F96AB9B9F91B" bold="true" box="[192,654,1671,1695]" pageId="20" pageNumber="19">
<emphasis id="92BDEA96FF92366FFF76F96ABAE8F91B" bold="true" box="[192,479,1671,1695]" italics="true" pageId="20" pageNumber="19">Liothorax alternatus</emphasis>
(Horn, 1870)
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<paragraph id="A0763684FF92366FFF76F959B9ABF948" blockId="20.[192,668,1716,1740]" box="[192,668,1716,1740]" pageId="20" pageNumber="19">
<bibRefCitation id="C4584B75FF92366FFF76F959BA95F948" author="Hendrickson, G. O." box="[192,418,1716,1740]" pageId="20" pageNumber="19" pagination="49 - 175" refId="ref32060" refString="Hendrickson, G. O. 1930. Studies on the insect fauna of Iowa prairies. Iowa State Journal of Science 4 (2): 49 - 175." type="journal article" year="1930">Hendrickson 1930</bibRefCitation>
; 
<bibRefCitation id="C4584B75FF92366FFE19F959B978F948" author="Gordon, R. D." box="[431,591,1716,1740]" pageId="20" pageNumber="19" pagination="157 - 167" refId="ref31565" refString="Gordon, R. D. 1977. Studies of the genus Aphodius of the United States and Canada (Coleoptera: Scarabaeidae). V. An evaluation of Horn's Group F. Proceedings of the Entomological Society of Washington 79 (2): 157 - 167." type="journal article" year="1977">Gordon 1977</bibRefCitation>
; ISIC.
</paragraph>
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<paragraph id="A0763684FF92366FFF76F936B917F977" blockId="20.[192,544,1755,1779]" box="[192,544,1755,1779]" pageId="20" pageNumber="19">Collection dates: April–May.</paragraph>
<paragraph id="A0763684FF92366FFF76F8EEBE58F8D0" blockId="20.[192,1440,1795,1876]" pageId="20" pageNumber="19">
<materialsCitation id="10A13CD9FF92366FFF76F8EEBA38F8BC" collectingDate="1926-04-16" collectionCode="ISIC" county="Story Co." location="Ames" municipality="Specimen" pageId="20" pageNumber="19" specimenCount="1" stateProvince="Iowa">
<collectingMunicipality id="4012ACFEFF92366FFF76F8EEBA00F89F" box="[192,311,1795,1819]" pageId="20" pageNumber="19">Specimen</collectingMunicipality>
label data: 
<location id="A516605FFF92366FFE79F8EEB920F89F" LSID="urn:lsid:plazi:treatment:28608792FF92366EFF76F96AB92DFF6B:A516605FFF92366FFE79F8EEB920F89F" box="[463,535,1795,1819]" county="Story Co." municipality="Specimen" name="Ames" pageId="20" pageNumber="19" stateProvince="Iowa">Ames</location>
, 
<collectingCounty id="49174E08FF92366FFD93F8EEB9ABF89F" box="[549,668,1795,1819]" pageId="20" pageNumber="19">Story Co.</collectingCounty>
, 
<collectingRegion id="620DF866FF92366FFD1CF8EEB9DEF89E" box="[682,745,1795,1818]" country="United States of America" name="Iowa" pageId="20" pageNumber="19">Iowa</collectingRegion>
, 
<date id="D4771044FF92366FFD41F8EEB891F89F" box="[759,934,1795,1819]" pageId="20" pageNumber="19" value="1926-04-16">
<collectingDate id="C433E9ACFF92366FFD41F8EEB891F89F" box="[759,934,1795,1819]" pageId="20" pageNumber="19" value="1926-04-16">16 April 1926</collectingDate>
</date>
, GH, determined O. L. 
<bibRefCitation id="C4584B75FF92366FFB67F8EEBE97F89F" author="Cartwright, O. L." box="[1233,1440,1795,1819]" pageId="20" pageNumber="19" pagination="413 - 462" refId="ref30815" refString="Cartwright, O. L. 1955. Scarab beetles of the genus Psammodius in the Western Hemisphere. Proceedings of the United States National Museum 104 (3344): 413 - 462." type="journal article" year="1955">Cartwright 1955</bibRefCitation>
(
<collectionCode id="C6D8AE41FF92366FFF7FF8CDBA31F8BC" box="[201,262,1824,1848]" pageId="20" pageNumber="19">ISIC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="10A13CD9FF92366FFEADF8CDBEABF8BC" box="[283,1436,1823,1848]" collectingDate="1930-04-03" collectionCode="ISIC" collectorName="H. E. Guthrie" county="Story Co." location="Ames" pageId="20" pageNumber="19" specimenCount="1" stateProvince="Iowa">
<location id="A516605FFF92366FFEADF8CDBA54F8BC" LSID="urn:lsid:plazi:treatment:28608792FF92366EFF76F96AB92DFF6B:A516605FFF92366FFEADF8CDBA54F8BC" box="[283,355,1824,1848]" county="Story Co." name="Ames" pageId="20" pageNumber="19" stateProvince="Iowa">Ames</location>
, 
<collectingCounty id="49174E08FF92366FFED9F8CDBAD3F8BC" box="[367,484,1824,1848]" pageId="20" pageNumber="19">Story Co.</collectingCounty>
, 
<collectingRegion id="620DF866FF92366FFE47F8CDB907F8B3" box="[497,560,1824,1847]" country="United States of America" name="Iowa" pageId="20" pageNumber="19">Iowa</collectingRegion>
, 
<date id="D4771044FF92366FFD8AF8CDB9EFF8BC" box="[572,728,1823,1848]" pageId="20" pageNumber="19" value="1930-04-03">
<collectingDate id="C433E9ACFF92366FFD8AF8CDB9EFF8BC" box="[572,728,1823,1848]" pageId="20" pageNumber="19" value="1930-04-03">3 April 1930</collectingDate>
</date>
, 
<collectorName id="0D3C5352FF92366FFD52F8CDB8A6F8B3" box="[740,913,1823,1847]" pageId="20" pageNumber="19">H. E. Guthrie</collectorName>
, determined O. L. 
<bibRefCitation id="C4584B75FF92366FFBCEF8F2BE72F8B3" author="Cartwright, O. L." box="[1144,1349,1823,1847]" pageId="20" pageNumber="19" pagination="413 - 462" refId="ref30815" refString="Cartwright, O. L. 1955. Scarab beetles of the genus Psammodius in the Western Hemisphere. Proceedings of the United States National Museum 104 (3344): 413 - 462." type="journal article" year="1955">Cartwright 1955</bibRefCitation>
(
<collectionCode id="C6D8AE41FF92366FFAE0F8CDBEA4F8BC" box="[1366,1427,1824,1848]" pageId="20" pageNumber="19">ISIC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="10A13CD9FF92366FFF77F8D1BE5BF8D0" box="[193,1388,1852,1876]" collectingDate="1931-04-29" collectionCode="ISIC" collectorName="W. E. Heston" county="Story Co." location="Ames" pageId="20" pageNumber="19" specimenCount="1" stateProvince="Iowa">
<location id="A516605FFF92366FFF77F8D1BA3EF8D0" LSID="urn:lsid:plazi:treatment:28608792FF92366EFF76F96AB92DFF6B:A516605FFF92366FFF77F8D1BA3EF8D0" box="[193,265,1852,1876]" county="Story Co." name="Ames" pageId="20" pageNumber="19" stateProvince="Iowa">Ames</location>
, 
<collectingCounty id="49174E08FF92366FFEA2F8D1BABEF8D0" box="[276,393,1852,1876]" pageId="20" pageNumber="19">Story Co.</collectingCounty>
, 
<collectingRegion id="620DF866FF92366FFE23F8D0BAE3F8D0" box="[405,468,1853,1876]" country="United States of America" name="Iowa" pageId="20" pageNumber="19">Iowa</collectingRegion>
, 
<date id="D4771044FF92366FFE69F8D1B9B1F8D0" box="[479,646,1852,1876]" pageId="20" pageNumber="19" value="1931-04-29">
<collectingDate id="C433E9ACFF92366FFE69F8D1B9B1F8D0" box="[479,646,1852,1876]" pageId="20" pageNumber="19" value="1931-04-29">29 April 1931</collectingDate>
</date>
[2], 
<collectorName id="0D3C5352FF92366FFD0BF8D0B854F8D0" box="[701,867,1853,1876]" pageId="20" pageNumber="19">W. E. Heston</collectorName>
, determined O. L. 
<bibRefCitation id="C4584B75FF92366FFBFEF8D1BE22F8D0" author="Cartwright, O. L." box="[1096,1301,1852,1876]" pageId="20" pageNumber="19" pagination="413 - 462" refId="ref30815" refString="Cartwright, O. L. 1955. Scarab beetles of the genus Psammodius in the Western Hemisphere. Proceedings of the United States National Museum 104 (3344): 413 - 462." type="journal article" year="1955">Cartwright 1955</bibRefCitation>
(
<collectionCode id="C6D8AE41FF92366FFA93F8D1BE54F8D0" box="[1317,1379,1852,1876]" pageId="20" pageNumber="19">ISIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A0763684FF92366FFF76F889B8BFF8F8" blockId="20.[192,904,1892,1916]" box="[192,904,1892,1916]" pageId="20" pageNumber="19">
County records (
<figureCitation id="38F22A01FF92366FFE3DF889B931F8F8" box="[395,518,1892,1916]" captionStart="Figures 58–65" captionStartId="68.[192,287,1847,1869]" captionTargetBox="[147,1439,217,1800]" captionTargetPageId="68" captionText="Figures 58–65. Distribution of Scarabaeoidea in Iowa.58) Geomyphilus insolitus (Brown, 1928). 59) Geomyphilus russeus (Brown, 1928).60) Labarrus cincticulus (Hope, 1847).61) Lechorodius terminalis (Say, 1823).62) Liothorax alternatus (Horn, 1870). 63) Melinopterus femoralis (Say, 1823). 64) Melinopterus prodromus (Brahm, 1790). 65)" figureDoi="http://doi.org/10.5281/zenodo.5353812" httpUri="https://zenodo.org/record/5353812/files/figure.png" pageId="20" pageNumber="19">Figure 62</figureCitation>
): Dickinson, Story, Winnebago.
</paragraph>
<paragraph id="A0763684FF93366EFF76FF3AB92DFF6B" blockId="21.[192,538,215,239]" box="[192,538,215,239]" pageId="21" pageNumber="20">
<taxonomicName id="67C94D07FF93366EFF76FF3AB92DFF6B" authority="Mulsant, 1842" authorityName="Mulsant" authorityYear="1842" box="[192,538,215,239]" class="Insecta" family="Aphodiidae" genus="Melinopterus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="92BDEA96FF93366EFF76FF3ABA56FF6B" box="[192,353,215,239]" italics="true" pageId="21" pageNumber="20">Melinopterus</emphasis>
Mulsant, 1842
</taxonomicName>
</paragraph>
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</treatment>
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