The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda
Author
Cowie, Robert H.
text
Zootaxa
2015
3940
1
1
92
journal article
10.11646/zootaxa.3940.1.1
5013eaff-bb37-4be3-a1ea-43c11d0cf5d1
1175-5326
288204
B5D45D1B-4B52-4F0B-8AF6-B587F8857475
Genus
PILA
Röding, 1798
POMUS Humphrey, 1797: 58
. Unavailable name: introduced in a work rejected for nomenclatural purposes (
Melville & Smith 1987: 318
).
Remarks
.
Nevill (1885: 8)
incorrectly considered
Nerita urceus
Müller, 1774
as the
type
species.
PILA
Röding, 1798
: 145
.
Type
species:
Helix ampullacea
Linnaeus, 1758
, by subsequent designation of
Dall (1904: 53)
.
AMPULLARIA
Lamarck, 1799
: 76
.
Type
species:
Helix ampullacea
Linnaeus, 1758
, by monotypy.
Remarks
.
Nerita urceus
Müller, 1774
, invalidly designated as the
type
species by
Dall (1904: 53)
.
AMPULLARIUS
Montfort, 1810
. Unjustified emendation of
Ampullaria
Lamarck, 1799
.
PACHYSTOMA
Guilding, 1828
: 536
.
Type
species:
Ampullaria globosa
Swainson, 1822
, by original designation (
Guilding 1828: 539
).
PACHYLABRA
Swainson, 1840
: 339
. Unjustified replacement name for
Pachystoma
Guilding, 1828
.
Type
species:
Ampullaria globosa
Swainson, 1822
.
Remarks
.
Swainson (1840: 339)
wrote “
Pachystoma
Guild., already used in Ichthyology”.
Pilsbry & Bequaert (1927: 172)
considered
Pachystoma
Guilding preoccupied by “
Pachystoma
Latreille, 1809
” (
Diptera
, not ichthyology), but the latter is correctly spelled
Pachystomus
(see also
Dall 1904
: 53;
Neave 1940
: 517;
Evenhuis & Thompson 1990
: 258;
Sabrosky 1999
: 229). The different spellings (
Pachystoma
and
Pachystomus
) mean that they are not homonyms (
Code
, Art. 56.2). There appear to have been no earlier introductions of
Pachystoma
(see
Neave 1940
: 517).
PACHYCHEILUS
Broderip, 1840
: 454
. Unjustified emendation of
Pachylabra
Swainson, 1840
.
Type
species:
Ampullaria globosa
Swainson, 1822
.
Remarks
.
Baker
(1930: 27)
considered the name an unjustified emendation and
Pilsbry (1956: 33–34)
considered it merely a suggestion of how to improve Swainson’s name.
POMUS
Gray, 1847
: 148
.
Type
species:
Helix ampullacea
Linnaeus, 1758
, by original designation (
Gray 1847: 148
).
PA
CH
Y
C H I L
U S
Philippi, 1851
: 7. Unjustified replacement name for
Pachylabra
Swainson, 1840
; junior homonym of
Pachychilus
Lea & Lea, 1851
(
Pachychilidae Fischer & Crosse
).
Type
species:
Ampullaria globosa
Swainson, 1822
.
TURBINICOLA
Annandale & Prashad, 1921
: 7
, 9.
Type
species—
Ampullaria nux
Reeve, 1856
, by original designation (
Annandale & Prashad 1921: 9
).
Remarks
. Synonym of
Pila
Röding
,
teste
Berthold (1991: 202)
.
The validity of the names
Pila
Röding
,
Ampullaria
Lamarck
and
Ampullarius
Montfort
were discussed by
Cowie (1997b)
and ruled on by ICZN (1999a).
+
adusta
Ampullaria adusta
Reeve, 1856a
: pl. 3, fig. 11.
Type
material
—
syntype
: NHMUK 20020690 (figured by
Brown (1980
: fig. 23c; 1994: fig. 23c); labelled “
Zanzibar
”).
Type
locality
—“probably from Borneo”.
Distribution
—endemic to
Zanzibar
(
Pain
1961
: 22, 1963: 152).
Remarks
.
Nevill (1885: 10)
gave the locality as “South
America
”. Confirmed as African (and hence to be placed in
Pila
) by
Alderson (1925
: 86; see also
Cowie & Thiengo 2003
: 80), although
Pilsbry & Bequaert (1927: 176)
expressed doubt about this, synonymizing it with the South American
sordida
Reeve. Both
adusta
and
pilula
Reeve
were first published on the same plate. G.B. Sowerby III (1910: 56) and
Alderson (1925: 86)
synonymized them, explicitly treating
adusta
as the senior synonym. Valid species,
teste
Kobelt (1912b: 55), who placed it in
Pachylabra
. Placed in
Pila
as a form of
ovata
Olivier
by
Brown (1980
: 45, 1994: 54), followed here.