Molecular phylogeny supports invalidation of Didelphodiplostomum and Pharyngostomoides (Digenea: Diplostomidae) and reveals a Tylodelphys from mammals Author Achatz, Tyler J Author Chermak, Taylor P Author Martens, Jakson R Author Woodyard, Ethan T Author Rosser, Thomas G Author Pulis, Eric E Author Weinstein, Sara B Author Mcallister, Chris T Author Kinsella, John M Author Tkach, Vasyl V text Zoological Journal of the Linnean Society 2022 2022-09-01 196 1 124 136 https://academic.oup.com/zoolinnean/article/196/1/124/6517659 journal article 133186 10.1093/zoolinnean/zlab114 bd2fe3a4-83a4-4b80-aeba-39a775595da2 0024-4082 PMC9416008 36051981 7031477 TYLODELPHYS DIESING, 1850 (AFTER NIEWIADOMSKA, 2002 , AMENDED) Diagnosis: Body linguiform, typically indistinctly bipartite; opisthosoma conical or ovoid. Anterior extremity of prosoma not distinctly trilobate; pseudosuckers present. Oral and ventral suckers and pharynx small or large; holdfast organ round or oval, with median slit for opening.Ovary ellipsoid or spherical, submedian, pretesticular, near anterior margin of opisthosoma. Vitellarium in prosoma and opisthosoma, extending anterior to the level of caecal bifurcation in prosoma and posterior to testes in opisthosoma in some species. Testes tandem, typically symmetrical with ventral concavities, forming horseshoe shape; anterior testis symmetrical or asymmetrical. Ejaculatory pouch present or absent. Ejaculatory duct joining uterus forming hermaphroditic duct. Genital cone small or absent, when present, hermaphroditic duct opening terminally. Copulatory bursa with subterminal or (rarely) terminal genital pore. In Accipitridae Vieillot , Ardeidae Leach , Didelphidae Gray and Podicipedidae . Cosmopolitan. Metacercariae of ‘diplostomulum’ type , in fishes or amphibians. Cercariae with four preacetabular penetration gland cells; flame-cell formula 2[(2+2)+(2+[2])] = 16. Type species Tylodelphys clavata ( von Nordmann, 1832 ) .